Blancan Carnivore Trigonictis 
17 
Florida do in fact represent a single individual, or at least reflect the true 
relative lengths of the elements, they are subequal in Trigonictis, as indi- 
cated also by the associated skeleton of T. cookii (cf. Table 3). In relative 
length, absolute size, proportions, and morphology, the Florida specimens 
compare most favorably with those of Pannonictis pilgrimi from the 
Villafranchian of Hungary described and illustrated by Mottl (1941), and 
further compared and the illustrations reproduced by Ficcarelli and 
Torre (1967). 
Measurements. — See Tables 1-3 for measurements and statistics. 
The distinction in size between Trigonictis macrodon and T. cookii is 
not clear-cut, yet all authors, including us, agree that the range in size is 
too great to be accommodated in a single species, or at least in a single 
penecontemporaneous population. We have followed Bjork (1970) in 
placement of the boundary, as reflected in his statistics presented in our 
Table 2, modified slightly by the addition of new specimens. Incidentally, 
there is an anomalous point for an Mj- assigned to T. cookii in Bjork’s 
figure 13, plotted at approximately 12.2 mm length and 4.8 mm width; 
this specimen does not appear in the statistics for either species. 
Gustafson (1978) presented a case for a chronocline of increasing size 
upward in the Hagerman beds. Presumably because the length of Mj- 
(12.0 mm) of the single specimen from the older White Bluffs fauna fell in 
the gap between the two species, he moved the boundary upward and 
referred the specimen to T. cookii. However, in the other variates pre- 
sented, the specimen falls within the observed range of T. macrodon, and 
we have referred it to the larger species. Gustafson’s measurements for 
UW 41527 include: length of P^, 6.5 mm; width of P^, 3.7; length of Mj-, 
12.0; width of Mj-, 5.3; depth of jaw below Mj-, 12.7 (cf. our Table 2). 
Bjork (1970) did not give depth of jaw below M^ for T. cookii, but two 
specimens from Hagerman in USNM measure 10.6 and 10.8 mm. The 
inferior border of the ramus is missing in the holotype (USNM 12606), 
but its depth could scarcely have exceeded 1 1 mm. 
Similarly, on the basis of size (length of Mj-, 12.5 mm), we regard 
F:AM 49163 from the Sand Draw local fauna as T. macrodon rather 
than T. cookii, as assigned by Hibbard (1972a: 109). His measurement of 
12.6 for “greatest length P^ - Mj-” is in error, and should be 18. 1. 
Based on our measurements from his figures, Shotwell’s (1970:82, fig. 
37J-N) specimens from the Grand View local fauna are correctly 
assigned, although the larger is in some dimensions slightly larger than 
other published specimens of Trigonictis, for example in length of Mj-, 15 
mm, and depth of jaw below Mj-, 17 mm. However, there is a fragment of 
a jaw, USNM 25027, from Hagerman, almost certainly representing 
Trigonictis, that would have been as large or even larger. 
