Appalachian Pseudanophthalmus 47 
0.76 wide, pronotum 0.76 long X 0.90 wide, elytra 2.30 long X 1.48 wide, 
antenna 2.70 long. 
Discussion. — This rare species is probably closest to P. holsingeri, 
with which it shares the same aedeagal pattern. It is remarkable that 
another species {P. paynei, tennesseensis group) exists in Moores Bridge 
Cave only 1.3 km south of Weaver Cave, on the same side of Clinch 
River, but P. paynei has not yet been encountered in Weaver Cave, nor 
has P. wallacei been taken in Moores Bridge Cave; the two species are 
potentially sympatric. From P. wallacei, P. paynei is distinguished by 
several pronotal characters: the hind angles are obtuse, the sides are 
convergent and not sinuate, and the disc bears two pairs of long setae. 
Pseudanophthalmus rotundatus Valentine 
Valentine 1932:271. 
For many years a very rare species, P. rotundatus was originally de- 
scribed from English Cave, also the type locality of P. engelhardti\ it is 
more robust than P. engelhardti, with more angular humeri, the prehu- 
meral borders perpendicular to the midline, deeper elytral striae, and a 
broader apical blade on the aedeagus. Jeannel (1949) reported the species 
from Parkeys Cave, Hancock County, Tennessee, farther up the Powell 
River valley. J. R. Holsinger and T. C. Kane recently took P. rotundatus 
and P. delicatus {hirsutus group) by trapping in Smith Cave, near Rose 
Hill, Lee County, Virginia. In July 1979, J. R. Holsinger, T. C. Barr, HI, 
and I collected 33 specimens of P. rotundatus in Subers Cave, 2.7 km 
east-northeast of Hopewell (Coleman Gap IVi Quadrangle, 36°34'32" X 
83‘^25'52"), Hancock County, Tennessee, in a locality intermediate 
between English Cave and Smith Cave. The beetles occurred beside a 
small stream, under rocks, sticks, and leaves. The range of this species 
thus extends along the Powell River valley for approximately 30 km, 
overlapping the range of P. delicatus at the northeast end and coexisting 
in English Cave with P. engelhardti and P. pallidus. 
Pseudanophthalmus sidus Barr 
Barr 1965:64. 
This rare species is known only from three specimens taken in Mere- 
dith Cave, at Shanghai Boat Dock, Campbell County, Tennessee, about 
45 km southwest of English Cave in the lower Powell River valley. It is 
smaller (3.6-4. 1 mm) than either P. engelhardti or P. rotundatus, subpar- 
allel, depressed, polished (microsculpture obsolescent at center of elytral 
disc); prehumeral borders perpendicular to midline, humeri angular; 
pronotum 0.8 as long as wide, hind angles very large and right. The 
aedeagus is shorter (0.51-0.55 mm) and less arcuate than that of P. rotun- 
