50 
Thomas C. Barr, Jr. 
In Tennessee Caverns, Byers Cave, and Johnsons Crook Cave P.fulleri 
coexists with P. digitus {hirsutus group). Related vicar species occur in a 
series of en echelon anticlinal valleys nearby: P.fastigatus in the valley of 
West Chickamauga Creek to the east, P. sequoyah in Wills Valley to the 
southwest, and P. nickajackensis still farther west in the Tennessee River 
valley. 
Pseudanophthalmus fulleri was the first of a series of closely similar, 
allopatric species to be discovered in southeastern Tennessee, northwest 
Georgia, north Alabama, and west-central Tennessee. This ^"fulleri series” 
includes all species of the engelhardti group in this southern area except 
the rather different P. loedingi. All species of the series possess a hatchet- 
shaped aedeagal apex with sharp ventral cusp, resembling the northern 
species P. engelhardti, P. rotundatus, P. sidus, and P. deceptivus in this 
respect. The elytral humeri are minutely serrulate in iht fulleri series, and 
occasional specimens have a single long seta on each side of the pronotal 
disc. The aedeagal apex in P. loedingi, in contrast, is deflexed, produced, 
attenuate, and finely but distinctly truncate; polytypic P. loedingi occu- 
pies caves of the Cumberland plateau north of the Tennessee River. 
South of the Tennessee River near and west of Guntersville, caves are 
occupied by polytypic P. meridionalis Valentine, a member of the fulleri 
series. 
Pseudanophthalmus fastigatus, new species 
Figs. 3, 8 
Etymology. — Latin fastigatus, “sloping down, tapered.” 
Diagnosis. — Resembles P.fulleri but more polished, humeri much 
sharper, greatest width of elytra at apical third, elytral apexes elongate- 
attenuate (not rounded), longitudinal striae deeper, intervals subconvex; 
aedeagus smaller, apical blade larger, wider, and more deflexed. 
Description. — Length 4.4-4. 5 mm (N = 3). Form moderately slender 
and elongate, subconvex; rufotestaceous, very polished, pubescent; ely- 
tral microsculpture finely and obsoletely transverse. Head 1.2 times longer 
than wide, labrum singly emarginate; last 2 segments of maxillary palp 
subequal; antenna two-thirds body length, segments rather thick. Prono- 
tum transverse, more than 0.8 as long as wide, greatest width in apical 
third behind level of anterior marginal setae (as in P.fulleri), width at 
apex only seven-eighths width at base, which is about 0.85 greatest width; 
anterior angles subdued, sides very shallowly sinuate in basal fifth, hind 
angles large and slightly obtuse, secondary angles of base broad, low, 
rounded. Elytra highly diagnostic: 1.6 times longer than wide, prehu- 
meral borders perpendicular to midline, humeri sharply angular, greatest 
elytral width in apical third (rather than at middle) with sides behind 
