Appalachian Pseudanophthalmus 
53 
apical groove very long and oblique, feebly bisinuate, running to 3rd stria 
via crosier well in advance of 7th umbilicate puncture; elytral apexes 
separately rounded. Aedeagus 0.57 mm long (paratype), closely similar to 
that of P.fulleri but considerably more arcuate. 
Type series. — Holotype male (American Museum of Natural History) 
and 6 paratypes, Ellis (=Sequoyah) Cave (AL 333), 3.3 km SW Sulphur 
Springs, T4S/R10E/S20, Dekalb Co., Alabama, 25 August 1965, S. B. 
Peck. Three paratypes, same cave and collector, 9 August 1965. 
Measurements (mm). — Holotype, total length 4.64, head 1.06 long X 
0.94 wide, pronotum 0.94 long X 1. 10 wide, elytra 2.73 long X 1.67 wide, 
antenna 3.08. 
Discussion. — This species occupies a cave at the northwest end of 
Wills Valley, west of the Wills anticline; it is thus geologically isolated 
from most other Dekalb County caves, which are east of the anticline 
axis and inhabited by P. alabamae. Pseudanophthalmus sequoyah coex- 
ists with P. assimilis, a smaller and rarer species belonging to the hirsutus 
group. 
Alabama caves are assigned a sequential number as they are added to 
the official list of caves in that state (Varnedoe 1973). For example, Ellis 
Cave is AL 333, and nearby Kudzu Cave, discovered and explored much 
later, is AL 734. 
Pseudanophthalmus steevesi, new species 
Fig. 1 1 
Etymology . — Patronymic honoring Mr. Harrison R. Steeves, Jr., 
Birmingham, Alabama. 
Diagnosis. — A member of the fulleri series, with subquadrate prono- 
tum and very shallow elytral striae; aedeagus closely resembling that of P. 
fulleri but more sharply recurved at apex. 
Description. — Length 4. 2-4.6, mean 4.4 ± SD 0.2 mm (N = 7). Form 
subparallel, depressed, pubescent, rufotestaceous; elytral microsculpture 
obsolescent-transverse, shining. Head slightly wider than long but basi- 
cally rounded; labrum with very low and wide median lobe; last 2 seg- 
ments of maxillary palp subequal; antenna two-thirds body length. 
Pronotum subquadrate, 0.8 as long as wide, widths at apex and base 
subequal and about 0.7 greatest width, which occurs at apical fifth; sides 
subparallel in basal sixth, anterior angles not prominent, hind angles 
large and more or less right (in some specimens slightly produced and 
very sharp), secondary basal angles prominent but rounded. Elytra sub- 
parallel, rather depressed, prehumeral borders slightly oblique and 
humeri less prominent than in other species of the fulleri series (especially 
in specimens from the type locality cave); longitudinal striae very shal- 
