54 
Thomas C. Barr, Jr. 
low, 3 or 4 inner striae complete and outer striae obsolescent, intervals 
flat; scutellar stria very short, vestigial; apical recurrent groove fairly 
long, oblique, feebly bisinuate or not, joining 3rd stria behind level of 7th 
umbilicate puncture. Aedeagus 0.55-0.59 mm long, about as in P.fulleri 
but not quite as arcuate, median lobe more slender, apex with ventral 
cusp more strongly recurved, parameres with 4 quite short apical setae. 
Type series. — Holotype male (American Museum of Natural History) 
and one paratype male, Randolph Cave, 1.7 km SW Blount Springs, 
Blount Co., Alabama (AL 414, T13S/ R3W/S12), 15 June 1967, Harri- 
son R. Steeves, Jr., and Thomas C. Barr, Jr. Female paratype, same cave, 
12 December 1965, S B. Peck. 
Measurements (mm). — Holotype, total length 4.25, head 0.73 long X 
0.76 wide, pronotum 0.78 long X 0.98 wide, elytra 2.39 long X 1.45 wide, 
antenna 2.84, aedeagus 0.59. 
Discussion. — This species has also been collected in Rickwood Cav- 
erns (AL 236; one male and one female, 24 November 1972, W. W. 
Torode), Bryant Cave (AL 355; female, 19 March 1966, S. B. Peck), and 
Horse Cave (AL 721; male, 11 April 1970, W. W. Torode). Rickwood 
Caverns and Bryant Cave are both in T13S/R3W near Randolph Cave, 
and Horse Cave is in T12S/R1W/S10, all three caves in Blount County. 
A total of 7 specimens of this species was examined, including the type 
series. 
tennesseensis group (new group) 
Aedeagus with apex not constricted before base, not arrow-shaped in 
dorsal view; apex more or less finely truncate and slightly knobbed; 
copulatory pieces as in engelhardti group, left piece large and spinulose. 
Length 3. 6-4. 9 mm, mean lengths 3.8-4. 1 mm. The group includes four 
depressed, pubescent species with 2 long setae each side of the pronotum 
disc. The species occupy caves in Roane, Anderson, Knox, and Union 
counties west and north of Knoxville, in the Appalachian valley of east 
Tennessee. Type species: P. tennesseensis Valentine. 
Distribution. — Roane, Anderson, Knox, Union cos., TN. 
Discussion. — All known species of the group are closely similar, 
reflecting relatively recent common ancestry, and they are best differen- 
tiated by aedeagal structure. Barriers separating the species include both 
structural/ stratigraphic ones and the Clinch and Powell rivers. From 
other small species with discal setae on the pronotum the species of the 
tennesseensis group are separated by the large left copulatory piece (very 
small in hirsutus group) and produced, apically knobbed aedeagus 
(bluntly rounded in hirsutus group and in P. pallidus, a small species of 
the jonesi group). 
