Appalachian Pseudanophthdlmus 
73 
Pseudanophtahlmus jonesi is known from three caves in Grassy Cove, 
a large polje along the plunging axis of the Sequatchie anticline in Cum- 
berland County, Tennessee: Grassy Cove (=Brady) Saltpeter Cave, the 
type locality; nearby Mill Cave; and Blowhole (=“The Gouffre”). Grassy 
Cove is a karst island in the interior of the Cumberland Plateau; caves in 
the Appalachian valley to the east are occupied by species of the engel- 
hardti and hirsutus groups. In common with P. scutilus and P. rogersae, 
which occur in another karst island. Pine Mountain, P. jonesi has the 
elytra widest behind the middle and the apical groove is not distinctly 
bisinuate. This is the only species of the group previously described. 
Valentine (1945) quite rightly noted its unusually attenuate form and 
speculated that it was a representative of a more widely distributed stock. 
Pseudanophthalmus scutilus, new species 
Figs. 40, 49 
Etymology . — Latin scutilus, “very lean.” 
Diagnosis. — Resembling P. jonesi in the indistinctly bisinuate apical 
groove and slightly ventricose elytra, but head proportionately larger, 
pronotum more cordiform, and elytral striae obsolete. 
Description. — Length 4.3-5. 4, mean 4.8 ± SD 0.3 mm (N = 1 1). Form 
elongate and slender, convex, with elongate appendages; color pale rufo- 
testaceous, shining but a little dull; elytral microsculpture finely trans- 
verse, a little confused. Head rather large, a fourth longer than wide, 
maxillary palps with penultimate segment about 0.7 as long as fourth 
segment, labrum with obsolete median lobe. Pronotum slender and cor- 
diform, only slightly wider than long (L/W 0.94-0.96), greatest width in 
apical fifth at level of anterior marginal setae, apex width slightly greater 
than base width and about 0.8 greatest width; disc convex, with 1 or 2 
long setae each side in addition to sparse micropubescence; anterior 
angles subdued, sides shallowly sinuate in basal seventh, hind angles 
sharp and a little more than right, secondary basal angles only suggested. 
Elytra elongate-oval, 1.8 times longer than wide, convex, deplanate 
around scutellum, prehumeral borders a little oblique to midline, humeri 
somewhat rounded, finely setose but not serrulate; discal pubescence 
short, fine, rather sparse; longitudinal striae obsolete, inner 3 striae 
barely visible, intervals flat, outer striae absent, no scutellar stria; apical 
groove rather short and deep, hardly bisinuate but with slight suggestion 
of anterior flexure, ending blindly at level of anterior apical puncture. 
Aedeagus 0.71-0.73 mm long, basal bulb not greatly enlarged, median lobe 
slender, moderately and evenly arcuate, apex finely attenuate but without 
knob, truncation, cusp, or other special modification; parameres short, 
with 4 terminal setae. 
