136 
Richard L. Hoffman 
As might be expected of a milliped that occurs in leaf litter in the 
spruce-fir forest of the Great Smoky Mountains as well as in limestone 
caves along the Clinch River 50 miles distant and 4000 feet lower, there is 
evident geographic variation in structure of the gonopods. This circum- 
stance was perceived and accounted by Shear (1972:184, Figs. 103, 104), 
although he did not go so far as to nomenclatorially formalize subspecific 
differentiation. 
Since the already manifested differences between the populations of 
the Tennessee Valley and the Great Smokies have now been further 
substantiated by the discovery of two new forms of fracta in the Cumber- 
latid Mountains and in the Nantahala Gorge, I think it desirable to 
introduce trinomial designations for these four isolates, and to confirm 
Dr. Shear’s suspicion that cottus might be a junior synonym of fracta. 
In connection with the citation of material in the following accounts, 
the standard acronymic abbreviations are used to indicate the location of 
specimens, thus: 
MC2 — Museum of Comparative Zoology 
NCSM — North Carolina State Museum 
RLH — Personal collection of the author 
USNM — National Museum of Natural History 
Pseudotremia Cope 1869:179. Shear 1971:162-193 (monograph). 
The general structure of the gonopods in Pseudotremia has been 
adequately described and figured by Shear. I wish here only to express an 
opinion about one anatomical feature and to suggest a slight refinement 
in nomenclature. 
As shown in his Figure 1 (of Pseudotremia hobbsi), the dominant 
elements of the gonopods (8th pair of legs) are the apparent coxal deriva- 
tives for which the term colpocoxite (Ribaut 1920) seems applicable. 
Immediately posterior to the pair of colpocoxites, and attached to them 
only by connective tissue (no direct musculature has been demonstrated) 
is a sclerotized, almost medially diastemmate structure (the “bifid lami- 
nae” of earlier workers) that Shear considers to represent the fused telop- 
odites of the gonopods. If this interpretation is correct, and I have no 
better alternative to suggest, perhaps the term “syntelopodite” might be 
employed in this genus (in the related taxon Cleidogona, the two struc- 
tures remain discrete or minimally attached at their bases). When the 
appendages of the genitalic complex are held in a resting position, the 
syntelopodite is tightly embraced basally by large coxal lobes of the 
posterior gonopods (9th pair of legs). 
The anterior (ventral) surface of the syntelopodite is generally modified 
in Pseudotremia by the development of a central basal structure, referred 
