Milliped Taxonomy and Distribution 
137 
to by Shear as the “telopodite process” and by him quite properly exploit- 
ed as a taxonomic character of considerable importance. A wide spec- 
trum of form is evident, from a low median convexity to enormously 
elongated and apically bifid blades projecting conspicuously between the 
colpocoxites. 
In a small group of apparently related species native to eastern Tennes- 
see, the syntelopodite process is somewhat enlarged distally, and apically 
produced into several projections of variable form and length. These 
species are P. cocytus Shear, P. cottus Shear, P. scrutorum Shear, and P. 
minos Shear, and the ensemble was referred to by Shear as the Cottus 
Group. I think he was quite correct in considering it “. . . one of the more 
coherent species assemblages in the genus Pseudotremia.'' With the pres- 
ent reinstatement of P.fracta as a senior synonym of cottus, it will be 
appropriate to likewise alter the group designation. 
Both geographically and anatomically, P. minos is somewhat disjunct 
from the others (known from northern Alabama, troglobitic rather than 
trogophilic, and with the syntelopodite process four- instead of three- 
pronged) and perhaps warrants assignment to a group of its own. Of the 
remainder, cocytus occurs in Anderson County, Tennessee, apparently 
sympatric with fracta but not yet collected at the same locality with it; 
scrutorum is known only from the type locality in Scott County, Tennes- 
see; and fracta occurs widely from the crest of the Great Smokies to caves 
and bluffs along the Clinch River. 
Pseudotremia fracta Chamberlin 
The original description of this taxon was based upon a single female 
from Gatlinburg, Tennessee, and mentioned in four brief sentences color- 
ation, size, number of ocelli, and form of the metatergal surface sculp- 
ture. Although the specimen was subsequently misplaced in the collection 
of the describer, and despite the brevity of the account, it was possible for 
Dr. Shear to suspect that the name fracta applied to an immature indi- 
vidual of the species that he (1972:1) named Pseudotremia cottus. Subse- 
quent to the demise of Dr. Chamberlin in 1968 his myriapod collection 
was transferred to the National Museum of Natural History and in 1977 
the diplopod material was placed in my hands for reorganization. Event- 
ually the type oi fracta was discovered and Shear’s prophesy fulfilled; it is 
indeed a female lacking one moult of maturity yet externally identical 
with specimens identified by Shear as P. cottus (which is, moreover, the 
only member of the genus so far known from the western slopes of the 
Great Smokies). 
The present decision to recognize taxonomically nameworthy popula- 
