34 
Marianne E. Filka and Rowland M. Shelley 
S. latior juveniles also were present, and the remaining species were 
represented only by scattered individuals. Five species, P. fasciculatus, A. 
magnum, Croatania catawba, P.c. incursus, and S. /an’or were more abun- 
dant as both adults and juveniles during July than at any other time. Im- 
matures of Nopoiulus minutus and Scytonotus granulatus also were most 
numerous in July, although juveniles of the former were taken in October 
as well. Species more common as adults in July than in April included 
Polyzonium strictum, N. americanus, and O. gracilis. Species less common 
as adults during July than in October or April included C. annulata and 
B. stricta. Three species — P. ectenes, P. branneri, and D. lea — were ab- 
sent from the July collections. Andrognathus corticarius was collected 
only in July. 
The greatest abundance and diversity of species occurred in October. 
Those most abundant as adults were Ptyoiulus ectenes, P. impressus, C. 
annulata, P. branneri, and A. erythropygos. Those most common in im- 
mature stages were P. ectenes, O. gracilis, P. strictum, N. americanus, and 
A. erythropygos. More juveniles of the last three species were encountered 
in October than in either of the other months. The first two species were 
moderately abundant as either adults or juveniles. Both Teniulus sp. and 
Cleidogona medialis were found only in October and as adults. Millipeds 
less common in the adult stage in October than in July included N. 
americanus, A. magnum, O. gracilis, C. catawba, P.c. incursus, and S. 
latior. 
The species/genus ratio (S/G) for all three months (1.04) was essen- 
tially unity (April and July S/G = 1.00, October S/G = 1.05), with the 
slightly higher fraction of October reflecting the presence of both species 
of Ptyoiulus. Seasonal changes in the faunal composition ratios (or- 
ders/families/genera/species, O/F/G/S) from April to October were 
more significant than changes in the S/G ratios. One more order, the 
same number of families, and one less genus and species occurred in July 
(8/12/16/16) than in April (7/12/17/17). The same number of orders, 
one more family, four more genera, and five more species occurred in 
October (8/13/20/21) than in July. Thus, the spring and summer faunas 
were less diverse than the October fauna. These fluctuations reflect varia- 
tions in times of maturation and breeding of the different species. 
The overall O/F/G/S ratio for the three months combined 
(9/16/23/24) showed one more order, three more families, three more 
genera, and three more species than occurred in any single month. This 
reflects the appearance and disappearance of species during the year, 
which is also indicated by the following seasonal trends. Five species — 
P. ectenes, C. annulata, P. branneri, B. stricta and D. lea — were more 
common in April than in July and again increased in abundance during 
October. A different five species — P. strictum, P. ectenes, C. catawba, 
P.c. incursus and S. latior — were more common in July than in either of 
the cooler months. Three species — P. strictum, P. ectenes, and P. im- 
pressus — were most abundant in October, and two — N. americanus and 
