150 
James D. Williams and Henry W. Robison 
Ozarka, new subgenus 
Type Species. — Etheostoma punctulatum {\g 2 iss\z)= Poecilichthys 
punctulatus Agassiz 1854 (original description, type locality Osage River, 
Missouri). 
Species Included in Ozarka.— £. punctulatum, E. cragini, E. pallididor- 
sum, E. boschungi, and E. trisella. Etheostoma trisella was originally 
placed in the subgenus Psychromaster by Bailey and Richards (1963) 
while the other four species have been referred to the subgenus 
Oligocephalus (Bailey and Gosline 1955; Collette 1965; Wall and 
Williams 1974). The subgenus name Ozarka is taken from the Ozark 
Mountains Physiographic Provice, which we believe to have been the 
center of dispersal for the subgenus. 
Diagnosis . — A subgenus of the genus Etheostoma of moderate to small 
size, adult males and females ranging from 40-70 mm standard length 
(SL). Cheeks and opercles usually naked or with embedded scales, except 
E. trisella which has scales on these structures. Lateral line incomplete, 
except complete in E. trisella. Lateral line scale rows 40-80, anterior por- 
tion not arched upward. Transverse scale rows 12-24, caudal peduncle 
scales 20-29. Branchiostegal membranes narrowly joined to overlapping; 
branchiostegal rays usually 6-6 and unsealed. Frenum present, broad and 
well developed. Preoperculomandibular canal pores usually 10, infraor- 
bital canal complete with 7 or 8 pores, supratemporal canal complete or 
interrupted. Preopercle entire. Caudal peduncle deep. Vertebrae 32-39, 
usually 34-37. Caudal vertebrae usually 18, 19, or 20. First interneural 
spine between the neural spines of third and fourth or fourth and fifth 
vertebrae. 
Dorsal spines VI-XII usually IX-XI; dorsal rays 10-15, usually 11- 
14; pectoral rays 9-14, usually 11-13; anal spines 2 except usually 1 in E. 
trisella; anal rays 6-10, usually 7-9. Males with breeding tubercles on 
scales of belly, around base of anal fin posteriorly to caudal peduncle, 
and on anal spines and rays and ventral surface of pelvic rays; tubercles 
absent in females. Sexual dichromatism pronounced in breeding adults, 
males brightly colored, females not. Breeding males with bold blue-black 
subocular bar, width one-third to two-thirds diameter of orbit. First dor- 
sal fin of breeding male with black marginal band (usually narrow an- 
teriorly, increasing in width posteriorly), submarginal orange to red- 
orange band, and blue-green to blue-black basal band. Submarginal and 
basal bands approximately equal in width. Second dorsal of breeding 
males without bright colors. Venter from pectoral and pelvic fin inser- 
tions to caudal peduncle suffused with varying concentrations of orange 
to red-orange pigment. Branchiostegal and gular region orange except in 
E. trisella. Genital papilla tubular in females, not long and tubular in 
males. Anus not surrounded by fleshy villi. 
