124—26 INDIAN TERTIARY AND POST-TERTIARY VERTEBRATA. 
These dimensions show that the anterior ‘ cannon-bone ’ of the hydaspithere 
forms another step from that of the helladothere in the direction of the giraffe. 
The head of the Indian hone shows a distinct remnant of the second metacarpal on 
its postero-internal angle, even more distinctly than that pointed out by Professor 
Gaudry in ILelladotherium while a peculiar projection on the outer side of 
the head appears to he the indication of a remnant of the fifth metacarpal. In the 
inferior portion of the hone there is a median septum dividing the medullary cavity 
into two tubes, hut superiorly these two tubes have coalesced. This septum indicates 
a close affinity with animals in which the metacarpals were not fused into a c can- 
non-bone/ 
Metatarsus . — We have no complete specimen of the metatarsus* A fragment 
of the condylar extremity belonging to the same individual as the metacarpus, shows 
the same dimensions across the condyles. There is a wide groove on the anterior 
surface, which is wanting in the metacarpus. The medullary cavity is divided into 
two divisions by a median septum. 
Proximal phalangeal. — The external proximal phalangeal hone associated with 
the metacarpus described above, is represented in figure 11 of plate XVII, of half the 
natural size. The hone is nearly as long as the corresponding hone of Sivatherium 
(plate XVII, figure 9), hut is much slenderer and slighter, approaching in this 
respect the giraffe. The proximal phalangeal of Helladotherium seems very similar 
to the present specimen. 
Calcaneum. — The last hone calling for any special notice is the calcaneum, of 
which a complete specimen from the Punjab is represented in figure 7 of plate XVII, 
of half the natural size. This hone is referred to the present species as being distinct 
from the calcaneum of Sivatherium , and as evidently belonging to- a large sized rumi- 
nant allied to the giraffe. It resembles the corresponding hone of the latter animal 
indeed, very closely, being mainly distinguished by its much larger size. It is 
readily distinguished from the calcaneum of Sivatherium , of which an imperfect 
specimen is represented in figure 12 of plate D of the ec Eauna Antiqua Sivalensis,” 
by its superior surface being broad and flattened for the greater part of its length 
in place of being produced into a sharp edge. In this respect the bone agrees with 
the calcaneum of the giraffe. The dimensions of this bone are compared below with 
those of the calcaneum of Sivatherium : — 
Sivatherium. 
Hydaspitherium. 
Total length 
8-6 
Projection of heel .... 
62 
5-5 
Breadth of calcaneal tuberosity 
29 
2-1 
Height „ „ „ . 
2-8 
2-0 
1 “ Les Enchainements du Monde Animal,” figure 147. 
