233—56 INDIAN TERTIARY AND POST-TERTIARY VERTEBRATA. 
punjabiensis , although faintly marked in E. sivalensis. On the external surface there 
is a very bold cingulum (partly damaged in the specimen) extending across both 
lobes of the blade : this cingulum is present in 71. sivalensis , but does not extend 
across the anterior lobe. 
In all the points in which this tooth differs from the first true molar of the other 
species of Eycenarctos , it approaches the more dog-like Dinocyon. In the first true 
molar of the latter 1 the antero-posterior shortening is carried to a still greater extent ; 
the inner cingulum is more developed, and the inner transverse ridge rendered less 
prominent. From this tooth there is but one step to Cephaloyale, 2 in which the 
cingulum is more widely separated from the internal ridge, which assumes a horse- 
shoe shape ; and thence to Ganis (woodcut, fig. 8), in which the ridge is divided into 
two distinct cusps. 
The last true molar of the specimen differs from the corresponding tooth of the 
two preceding species of Eycenarctos by the oblique truncation of its postero-external 
angle, and the entire absence of any backward prolongation of the postero-internal 
angle. In consequence of this the hinder external lobe becomes less distinct and is 
placed more internally. In respect of the obliquity of the outer ridge, E. punjabiensis 
comes nearer to the specimen than the other species. The internal ridge is curved, 
and has a well-marked cingulum. The absence of any trace of a hind talon to this 
tooth is one step further away from TJrsus than is made by the other species of 
Eycenarctos : the oblique position of the line of the external lobes is a dog-like 
character, displayed in the last molar of Dinocyon : in that genus, however, the last 
molar is smaller antero-posteriorly than the penultimate, although, as in Cephaloyale , 
this character is not so strongly marked as in Ganis. The last molar of the present 
specimen is, therefore, intermediate between the corresponding tooth of the other 
species of Eycenarctos and Dinocyon . 
In the following table the dimensions of the specimen under consideration are 
compared with those of Eycenarctos sivalensis and 77. punjabiensis , and Dinocyon thenardi 
and D. hemicyon : — 
Length of last three cheek-teeth 
,, ,, pm. 4 
Width ,, ,, ,, (at middle) . 
Length , , m. 1 (outer ridge) 
,, ,, „ ,, (inner „ ) 
Width „ „ „ 
Length,, ,, 2 (inner ridge) . 
,, ,, ,, ,, (outer ,, ) . 
Width ,, ,, ,, (anteriorly) 
,, ,, ,, ,, (posteriorly) 
•£ 
§ 
^ 3 •48 
1-25 1-19 
0- 88 0-94 
1- 18 1-2 
M2 1-18 
1-08 1-08 
Ml 
0- 96 
1- 05 
0-9 
I 
3-44 
1-3 
0-78 
1-2 
0- 98 
1- 05 
11 
1-0 
1-2 
1-1 
1 ! J 
m O O 
3-1 
1-09 
0-81 
1-06 .. 1-05 .. 1 09 
0- 92 .. 0-75 .. 0-95 
1- 06 .. M4 .. 1-38 
1-0 .. 0-77 .. 1-02 
1-05 .. 0-77 .. 1-05 
M3 .. 1-04 .. 1-45 
0-97 .. 0-95 .. 1-15 
i Gervais, op. cit., pl.-LXXXI 
2 Filhol, op. cit., pi. II., fig. 4. 
Filhol, op. cit., pi. III., fig. 11. 
