245—68 INDIAN TERTIARY AND POST-TERTIARY YERTEBRATA. 
hind talon large : pm. 4 has its internal tubercle large, and placed further back than 
in Ganis , thus indicating affinity with Dinocyon hemicyon : m. 2 is relatively large : m. 1 
is almost exactly like that of Ganis. The base of the skull and the limb-bones 
present considerable resemblances to those of Amphicyon. 
The genus Vulpavus has been formed by Prof. Marsh 1 on the evidence of 
some upper molars from the eocene of Wyoming : these teeth are said to differ from 
those of existing foxes, by their relatively larger transverse diameter. 
Of the remaining genera, Brachycyon , 2 Filhol, from the Quercy phosphorites, is 
a meionocreodont form, only known by the mandible, and distinguished by the 
absence of pm. 1 ; the number of the lower cheek-teeth being Pm. 7, M. 7. It is 
considered to be closely allied to Amphicyon. 
Temnocyon , 3 Cope, from the miocene of North America, has Pm. f , M. § : this 
genus is said to be distinguished from Ganis by the hind talon of mTi being 
trenchant, in place of cup-sliaped, and by the presence of an epitrochlear foramen 
to the humerus : in the former character it agrees with Icticyon, and in the latter with 
Amphicyon and the bears. 4 
Enhydrocyon , 5 Cope, from the same formation is distinguished from Temnocyon 
by the absence of the first premolars, and perhaps of m. 3 ; the formula of the 
cheek-teeth being Pm. f , M. ^ : it agrees with Temnocyon in the trenchant form 
of the hind talon of m. 1. 
In Uycenocyon , 6 Cope, also from the miocene of North America, the formula of 
the cheek-teeth is Pm. M. ^L. It is somewhat doubtful if this genus belongs to the 
Ganince ; if it does, its dentition is numerically more specialized than that of any 
other genus ; Icticyon coming nearest in this respect. 
Palceocyon? Lund, from the pleistocene of South America, has Pm. f, M. f , and 
agrees with Icticyon in the absence of the tubercle of pm. 4 , and the inner cusp of the 
blade of m. 1 : it appears to be megalocreodont. 
Speothos, Lund, of the same deposits, is considered by Prof. Huxley 8 to be 
probably the same as Icticyon. 
Ly corns? Bourg 1 , is a megalocreodont wolf from the caverns of France, of 
which only the mandible and lower dentition are known : it agrees with Enhydrocyon 
and Brachycyon in the suppression of pm. I, - the number of the lower cheek-teeth 
being Pm. 7, M. 7; but differs from the former in the cup-sliaped talon of m. 1. In 
the opinion of the present writer the mere absence of pm. 1 is not sufficient to justify 
generic distinction from Ganis. 
1 “American Journal of Science,” 3rd ser., vol. II., p. 124. The genus Dromocyon, Marsh (Ibid), vol. XII., p. 403, is 
allied to Uymnodon. 
2 Filhol, “ Phosphorites du Quercy,” figs. 27-9. 3 Cope, op. cit., p. 179. 
4 M. Filhol (“ Notes sur quelques Mammiferes Fossiles, etc.,” p. 82) quotes Prof. Cope to the effect that the ursine post* 
parietal foramen is present in this genus : in the passage quoted, however, the presence of this foramen is denied by Prof. Cope. 
3 Cope, op. cit., p. 178. 6 Ibid, p. 181. 7 Huxley, op. cit., pp. 279-80. lalcccyon, Blain .=Arclocyon. 
8 Ibid, p. 280. 9 Bourguignat, ‘Ann. d. Sci. Geol.,’ vol. VI., pi. XVIII. 
