SIWALIK AND NARBADA CARNIVORA. 
69—246 
The genus E llocyon, Aym., may perhaps be referred to the Viv err idee. Arctocyon , 
Blain., is too imperfectly known for its position to be accurately determined; Prof. 
Cope makes it the type of a family of his order Creodonta. Hemicyon , Lartet, is 
provisionally referred to Dinocyon ; and Pseudocyon , Lartet, to Amphicyon. Hydrocyon , 
Lartet, is said to have a dentition intermediate between that of the dogs and the 
Genus I. : AMPHICYON, 1 Lartet. 
Syn. Agnotheriwn, Kaup (in parte). Cynelos, Jourd. (?) Pseudocyon , Lartet. 
Dentition . — As this genus is more nearly allied to the bears than Oanis, it is 
placed first : many of its dental characters have already been mentioned, and it will, 
therefore, suffice to indicate in what respects the individual hinder cheek-teeth differ 
from those of Canis. According to M. Filliol, 2 pmT3 may or may not have a posterior 
talon-Cusp ; the latter being present in all species of Canis except C. palceolycos. In 
m. 1 the inner cusp of the blade is larger and placed more forward than in Canis , 
although some of the meionocreodont forms of the latter make a near approach in 
this respect : this tooth in AmpMcyon is in fact intermediate between that of Cynodictis 
and Canis. In pm. 4 the inner tubercle is larger than in Canis, the whole tooth being 
intermediate between the carnassials of Canis and Cynodictis. The two last lower 
true molars are almost indistinguishable from those of Canis. In m. 1 the two outer 
lobes are generally placed more internally than in Canis , forming two isolated 
pyramids on the crown, instead of being continuous with the outer wall as in the 
latter : A. ambiguus is, however, intermediate in this respect : in typical species this 
tooth approaches Cynodictis. This tooth is also more developed transversely than in 
Canis ; its cingulum embracing more of the inner half of the crown (though there is 
a certain amount of variation in this respect between different individuals of C. 
lupus) : and the two inner tubercles, in place of being distinct as in Canis , united into 
a crescent placed internally to the cingulum. 
Distribution. — According to Prof. Gaudry 3 the earliest appearance of the genus 
in Europe is in the middle eocene [Stage des sables de Beauchamp) ; and it survived till 
the middle miocene of Sansan. It is represented in the miocene of North America, 
and in the Siwaliks of India. 
Number of species .- — The following list comprizes the best determined of the 
named species, exclusive of the Siwalik form, which is described below. Owing to 
the confusion in the synonomy of the European species, it is possible that this list 
may not be in all respects perfectly correct. 
1. Amphicyon ambiguus, 4 Filhol. Quercy phosphorites. 
M. 3 very small, and may be absent : pm 4 sometimes very large : m. 1 more canoid than 
in any other species : the species was about equal in size to C. lupus. 
1 The name Amphicyon was applied by Lartet in 1837 (‘ Compt. Bend.,’ vol. V., p. 424), and that of Agnotlierium by 
Kaup in 1833 (“ Oss. foss. d. Musee d. Darmstadt”) : the latter has, therefore, the priority, but the former has gained 
universal acceptation. 
2 “ Notes sur quelques Mammiferes Fossiles, etc.,” pp. 78-81. 
3 “ Les Enchainements du Monde Animal, etc.— Mammiferes Tertiaires,” p. 4. 
4 “ Phosphorites du Quercy,” figs. 22-6, 41-3. 
