SIWALIK AND NARBADA CARNIVORA. 
133—310 
Other unnamed specimens . — In the course of tlie preceding descriptions nearly all 
the accessible remains of Siwalik liysenas have been, at all events provisionally, 
named. There are, however, in the British Museum (Nos. 16,583; 39,138; and 
39,179) three specimens of the muzzles of Siwalik hyaenas 1 which are not specifically 
determinable. The same collection also contains a few undetermined detached 
teeth ; and there are some fragmentary mandibles in the Ipswich Museum, probably 
belonging either to H. colvini or II. felina. It is noteworthy that, with the exception 
of a few metapodials and phalangeals, no limb-bones have been obtained from 
the Siwaliks that can be referred to liysenas ; — indicating the great rarity of 
carnivorous remains. 
Mutual Affinities of the Species of Hyaena. 
As the hyaenas from the Siwaliks are so strongly represented in species it will 
be advisable in conclusion to cast a brief glance at their general relations to the 
other species, and the mutual affinities of the whole number. For this purpose the 
Siwalik species have been incorporated below in the list given on page 278 ; the 
order of arrangement following as far as possible the relative specialization of the 
dentition ; the most highly specialized forms being placed at the top, and doubtful 
species indicated by an asterisk: — 
t 
A ^ 
3rd lobe, pm.4 
m.l 
pm. 1 
cusp 
talon 
m .2 
H. croeuta (inner cusp absent in mm. 4) 
*H. perrieri ...... 
large 
small 
absent 
absent 
small 
absent 
H. colvini (pm. 1 absent) .... 
( generally i 
' \ smaller 1 
| medium, 
[ to large 
», 
„ 
„ 
H. felina (pm. 1 sometimes absent) 
smaller 
small 
,, 
„ 
larger 
H. eximia (tubercle of pm.4 very small, P m - ■ 
l in one 
instance wanting) .... 
• 
large 
present 
,, 
large 
>» 
H. brevirostris . . 
H. grseca (inner cusp present in mm. 4 ) 
• • 
very large 
P 
present 
| 
” 
present 
H. brunnea (inner cusp of md small) 
small 
large 
absent 
present 
large 
absent 
*H. arvernensis (inner cusp of mTl large) 
,, 
„ 
,, 
larger 
„ 
H. striata ,, ,, ,, ,, ,, 
H. sivalensis (inner cusp of md small : pr 
emolars 
„ . 
” 
» 
” 
” 
somewhat elongated) 
A, f 
very large 
„ 
„ 
very large j 
' sometimes 
present 
H. macrostoma (premolars much elongated) 
,, ,, 
present 
present ? 
H ?■ I 
? 
H chseretis ,, ,, ,, 
• 
? 
„ 
present 
very large 
'? 
It will be seen from this table that, although the species cannot, in all respects, 
be arranged in a regular series, the specialization of the dentition advances 
on the following lines : — first, in the gradual proportionate increase of the 
third lobe of pm. 4 • second, in the decrease in size of m. 1 ; third, in the disappearance 
of pm. I and m. 2 : fourth, in the diminution of the talon, and the diminution and 
eventually disappearance of the inner cusp of m. 1 ; which, however, lingers longer 
in mm. 4 : fifth, in the proportionately increasing width of the premolars. The 
carnassials also become larger in proportion to the other teeth ( megalocreodontism ). 
G 2 
i “ F.A.S.,” supl. plate L., figs 1, la ; 2, 2a. 2b ; 3, 3a, 3b. 
