SIWALIK AND NARBADA CARNIVORA. 
151—328 
masseteric fossa is, as in the fossils, more sharply defined than in most living cats ; 
while the lower incisors project slightly in front of the canine, although not to such 
an extent as in the fossils : pm. 3 has, however, large fore-and-aft talons, which are 
wanting in the latter. 
The present jaws have their canines and diastemse too small to permit of their 
belonging to F. cristata } The mandible of F. arvernensis, Cr. and Job., 2 presents a 
strong resemblance to the present specimens ; having cheek-teeth of the size of a 
small tiger, but the canine smaller ; the incisors proj ecting beyond the latter ; a small 
diastema ; and a sharply defined masseteric fossa. The diastema is, however, not 
quite so short, while the talons of pm” 3 are more developed than in the fossils under 
consideration ; these points being probably indicative of specific distinctness. 3 
Apparently the only other species to which the present specimens could belong is the 
little known F. hr achy stoma* ; but the canine of that form is considerably stouter. 
From the foregoing comparisons it is evident that the present specimens indicate 
the existence in the Siwalik epoch of a feline carnivore nearly as large as a small 
tiger, but readily distinguished by its shorter jaws and smaller canines. That it 
presented a very strong resemblance to the living genus Cyncelurus is certain ; but in 
the face of certain differences in the form of pmT3, and in the absence of the 
characteristic pm. 4 , it would be hazardous to say that it belonged to that genus ; and 
it is therefore safer to refer it provisionally to the typical genus Felis ; especially 
since it is not improbable that were the skull of this interesting form known it might 
exhibit characters common both to Felis and Cyncelurus , and, in conjunction with F m 
arvernensis , remove the present distinctions between the two. The specific name 
brachygnathus may be appropriately applied to the Siwalik fossil. 
Species 3 : Felis (non. det. ; allied to F. pardus). 
Mandible. — In figures 4, 4a, of plate XLIII. there is represented the hinder 
portion of the left ramus of the mandible of another Siwalik Felis , in the British 
Museum, apparently indicating a third species of the genus. The specimen shows 
the three cheek-teeth, of which the two last are considerably damaged ; but still 
sufficiently perfect to show that the specimen belonged to a true Felis. The jaw is 
considerably smaller than that of F. brachygnathus , and cannot, therefore, belong to 
F. cristata. It differs from the former in the less well-defined masseteric fossa, in 
the more distinct talons and lower crown of pm. 3 ; and also in the absence of any 
sign of the symphysis, which in that species commences a short distance in front of 
the latter tooth. The diastema of the present specimen was, therefore, probably 
1 The reader may convince himself of the impossibility of these jaws belonging to F. cristata by making a tracing of 
one of the figures and placing it in its proper position on plate XLII. 
2 De Blainville, op. cit., pi. XVI. 
3 De Blainville was inclined to refer F. arvernensis to F. pardus : the diastema is, however, relatively shorter, and the 
crown of pm73 taller. 
i Bourguignat, op. cit., fig. 6. 
