SIWALIK AND NARBADA CARNIVORA. 
155—332 
Species 6 : Felis (?) sp., non. det. 
Upper canine. — In figure 5 of plate XLV. there is represented the left upper 
canine of a small carnivore ; which from its flattened inner surface must almost 
certainly have belonged to some cat-like animal. The summit of the crown has 
been broken away. The flattened inner surface is bounded by distinct vertical 
ridges ; but the vertical grooves present on both sides of the upper canines of most 
species of Felis, are absent. These grooves are, however, wanting on the inner side 
of the canines of F. isabellina ; generally on both sides of those of F. caracal 1 ; and 
always in Cyncelurus 2 ; so that on this account there is no reason why the specimen 
si' oul d not belong to Felis. 
Of the known Siwalik cat-like animals the only ones to which the present 
specimen could possibly belong are (1st) the liysenoid Lepthycena ; (2nd) FEluropsis ; 
(3rd) JElurogale ; (4tli) Felis, sp. 3 ; (5th) Felis, sp. 4 ; and (6th) F. subhimalayana. 
Of these, Nos. 2, 3, and 4 had canines considerably larger than the present specimen ; 
while in No. 6 the canine is much smaller. Judging from the depth of the jaw of 
No. 1, it is probable that the canine of that species was larger than the present 
tooth ; and since the canine of the isabelline lynx is also considerably larger, it is 
probable that the same was the case with the allied No. 5. It, therefore, seems that 
the tooth under consideration indicates another species of feline animal, which may 
be provisionally referred to the type genus. This species was probably about equal 
in size to F. ssrval ; or intermediate between the fourth Siwalik species and F. 
subhimalayana. The antero-posterior diameter of the base of the crown of the canine 
is 032 ; and the transverse 026. 
Genus IV. : MACH^RODUS , 3 Kaup. 
Synonyms. Agnotherium, Kaup [in parte). Cultridens , Cr. Drepanodon, Auct. 
Meganihereon, Auct. Smilodon, Lund. Steneoclon , Cr. Trucifelis, 
Leidy ( teste Cope). 
Distribution and number of species. — The most important dental and cranial 
characters of the genus having been already mentioned, it only remains to consider 
its distribution and the number of species. In time it apparently commenced in 
the upper eocene and continued down to the pleistocene period : the miocene 
and (probably eocene) species had smaller canines than those of the later periods . 4 
l Busk, ‘Trans. Zool. Soc. Lon.,’ vol. X., p. 83. 2 Owen, “ Odontography,” p. 487. 
3 The name Machmrodu « [Mnchairodus) was given hy Kaup in 1833 (“ 0>s. Foss. Mu*. d. Darmstadt ”), on the assumption 
that the remains on which it was founded belonged to a reptile. Steneodon was published in the same year (E. Geof., ‘ J> '<y. 
Eneychptsd.,' 1833): and Cultridens apparently in 1837 (Huot, “ Xonreau Coins elementaire de Giilogu," Paris). Drepanodon 
and Mrgauthe< eon, which are usually cited respectively as Nesti and Croizet," were assigned merely as specific names and have 
no claim to stand for the genus ( Vide Gaudry, “ An. Foss, et Geol de I'Atliqxe," pp. 106, 108). The question of priority lies, 
therefore, entirely between Machoerodus and Steneodon ; this has, however, been overriden hy the universal acceptance of the 
former. 
4 The allied, hut highly specialized, eocene genus Ettsmilus is apparently a very remarkable exception to this generalization. 
