345—168 INDIAN TERTIARY AND POST-TERTIARY VERTEBRATA. 
be the third incisor. The fang of the tooth is unusually large, and considerably 
bent ; the latter character being not improbably due to some lesion of the j aw of the 
animal to which it belonged. From the very large size of the specimen it may be 
pretty safely referred to the present species. 
The specimen is of some importance, as it readily distinguishes M. palmndicus 
from M. latidens ; in which the corresponding tooth 1 has distinct cusps at the base of 
the inner sides of the fore-and-aft trenchant ridges, which are totally wanting in the 
Siwalik tooth. 
Affinities . — The foregoing comparisons indicate the distinctness of the present 
form from any sufficiently described species ; and also that it was more nearly allied 
to the other Siwalik species and the American forms than to any of the European 
species, although in the structure of the mandible and pm. 3 it presents certain 
indications of affinity with the latter; thus confirming the conclusions previously arrived 
at, as to the inadvisability of generically separating the American and European 
forms. 
The relationship of the Indian and American maclnerodonts is a very remarkable 
one ; and the more so since several genera of Siwalik carnivores (e.g. Hycena and 
Mellivora) are essentially Old World forms. This relationship probably indicates a 
line of passage for the Siwalik and American forms through the regions to the west- 
ward of China. 
Distribution. — Remains of the present species have been obtained from the 
typical Siwalik Hills to the Punjab. 
Limb-bones of Felines. 
General. — It has been already observed that in the reference of the fossil limb- 
bones of the Delidce to their respective species, relative size is in the main the only 
guide ; and since in the case of the Siwalik representatives of the family there is 
frequently more than one species of the same approximate size, this guide is little 
better than useless. It has, therefore, been found advisable only to describe some of 
the more perfect of these remains, taking them in their serial order ; and making 
suggestions as to the species to which they may possibly belong. 
Humerus. — In figure 11 of plate XLIV. there is represented the anterior surface 
of the distal extremity of the right humerus of a large feline in the Siwalik 
collection of the British Museum (No. 37,146); in which there is another very 
similar specimen (No. 37,142), broken off at the supra-condylar foramen (c.f). The 
dimensions of these two specimens are compared with those of a humerus of the 
cave-lion, from Clacton (B. M., No. 28,014); viz . : — 
Cave-lion. Siwalik feline. 
Width across condyles 3 - 8 
,, ,, articular surfaces 2- 78 
,, of shaft above supra-condylar foramen . . . 2 -3 
37,146. 37,142. 
3-5 3-9 
2-35 2-75 
2-1 
l “ Brit. Foss. Mamjn. and Birds,” fig. 70, p. 182. 
