Genus Scytonotus 
5 
can continent. Gervais (1847) recognized the original combination 
a,s did Wood (1865), Bollman (1893), and Kenyon (1893a). Koch 
(1847) erected Scytonotus for three new species, all now synonyms 
of S. granulatus : S. scabricollis, from an unspecified locality in 
North America, and S. laevicollis and nodulosus, both from Penn- 
sylvania; Koch (1863) repeated these accounts. Sager (1856) pro- 
posed Stenonia hispida for an individual from Ann Arbor, Michigan, 
and Wood (1865) erected Polydesmus setiger for another specimen 
from Pennsylvania. Both names are additional synonyms of S. 
granulatus as is S. cavernarum , proposed by Bollman (1887) for a 
form from Indiana. Cook and Cook (1894) synthesized the earlier 
efforts and excluded five species that had been assigned to Scytonotus 
from Venezuela, New Zealand, Mexico, and the tropics. Under S. 
granulatus or a synonym, this common, eastern milliped has been 
recorded numerous times, and I have endeavored to list all refer- 
ences beside the appropriate names in the synonymy. 
The second eastern species, S. virginicus (Loomis), type spe- 
cies of the genus Lasiolathus (Loomis 1943), was proposed for an 
immature specimen from Thornton Gap, Page/Rappahannock coun- 
ties, Virginia. Hoffman (1947) synonymized the name with S. 
granulatus before collecting an adult male topotype and realizing 
(Hoffman 1950a) that S. virginicus is a distinct species. In his 
latter work, he gave the range as being the Blue Ridge Province 
south to Linville Falls, North Carolina. Chamberlin and Hoffman 
(1958) included S. virginicus in their checklist. Hoffman (19626) 
altered the range by stating that it was unknown south of the Roanoke 
River and proposed the final two eastern species, S. australis and 
michauxi, from north Georgia and western North Carolina/eastern 
Tennessee, respectively. 
Scytonotus was first recorded from western North America by 
Cook (1904), 83 years after Say (1821) described P. granulatus 
from the east. He reported juveniles and females of an unidentified 
species from Yakutat Bay, Sitka, and Juneau, Alaska, and illustrated 
a male from an unknown locality that he recognized as a new 
species. Unfortunately, Cook misplaced the specimen, so Chamberlin 
(1911) is credited with the authorship of S. bergrothi based on 
specimens from Bremerton, Kitsap County, Washington. Chamberlin 
(1910) described S. piger and the synonym, Polydesmus amandus , 
from Mill Creek Canyon, Salt Lake County, Utah, and 10 years 
later (Chamberlin 1920) added S. columbianus from the “Columbia 
Valley,” British Columbia, Canada. The precise locality is unknown, 
but he was probably referring to the valley of the Columbia River, 
which arises in this Canadian province. Chamberlin (1925) proposed 
