Genus Scytonotus 
13 
Charlotte Islands, where it never has been encountered. In the inte- 
rior, Scytonotus occurs in the Columbia River Valley and the Rocky 
and Selkirk mountains, and associated ranges, from Revelstoke and 
Yoho National Parks, British Columbia, to southeastern Washington, 
western Montana, and the southern extremity of the Idaho Pan- 
handle near the Salmon River. It is also known in the Wasatch and 
Teton mountains, and associated ranges, in western Wyoming, east- 
ern Idaho, and northern Utah south to Salt Lake County. 
Species — Nine, three in eastern North America, one divided 
into two geographic races, and six in the west. The species are 
arranged into three lineages or species groups, which are named for 
the oldest component. 
Key to Species of Scytonotus , based on adult males 
1. Distal lamina of gonopodal endomerite with 2 or 3 distal 
teeth (Figs. 5-8, 21-28) 2 
Distal lamina otherwise 6 
2. Medial lamina with 2 distinct teeth (Figs. 23, 25); 
Warren County, Virginia, to Sevier County, Tennessee, 
and Swain County, North Carolina 
virginicus (Loomis) 
Medial lamina with 1 or no teeth, with or without a 
rounded, basal lobe or flange 3 
3. Distal lamina with 3 teeth, the apical one much smaller 
and preceded by 2 larger, subequal teeth (Fig. 21); Ontario 
and Quebec to South Carolina, Tennessee, Arkansas, 
and Kansas granulatus (Say) 
Distal lamina with 2 subequal teeth 4 
4. Medial lamina with distinct distal tooth (Fig. 27); Bun- 
combe County, North Carolina, and Sevier County, 
Tennessee, to Oconee County, South Carolina, and Daw- 
son County, Georgia australis Hoffman 
Medial lamina with variable basal lobe but without distal 
tooth 5 
5. Endomerite much shorter than tibiotarsus; medial lamina 
a short, narrow flange, not overhanging inner margin 
of endomerite (Fig. 7); interior of British Columbia and 
northeastern Washington columbianus Chamberlin 
