Genus Scytonotas 
15 
the inner margin of the distal lamina. The medial lamina varies in 
length and is typically expanded into a basal lobe or flange, except 
on the Blue Ridge endemics, which have one or two distinct teeth. 
The lateral lamina is long, slender, and generally unmodified, ex- 
cept for S. granulatus and virginicus, in which the structure also 
has one or two teeth. Finally, the endomerite is shorter than or 
subequal in length to the tibiotarsus. The Blue Ridge species and 
S. columbianus have normal paranota on all segments in females, 
and males lack the tibial lobes. However, because all other conge- 
ners possess reduced paranota in females and lobes in males (Table 
1), I regard the conditions in these three species as apomorphies 
representing secondary simplification rather than plesiomorphies, and 
the similarity between S. columbianus and S. virginicus/ australis 
represents convergence. Spatial distributions tend to be allopatric 
and parapatric, with only minimal sympatry between S. virginicus 
michauxi and australis, in the Great Smoky Mountains, and be- 
tween S. piger and columbianus, in northwestern Washington. Rela- 
tionships among the components are hypothesized as columbianus + 
( piger + (granulatus + (virginicus + australis ))) (Fig. 34). The Blue 
Ridge Province seems to be a secondary center of evolution after 
the presumptive primary source area in the Cascade Mountains. The 
western representatives of the granulatus lineage are detailed below; 
the eastern species are characterized at the conclusion of the de- 
scriptive section. 
Components — granulatus (Say), piger Chamberlin, columbianus 
Chamberlin, virginicus (Loomis) (v. virginicus, v. michauxi Hoff- 
man), australis Hoffman. 
Scytonotus piger Chamberlin 
Figs. 2-6 
Scytonotus piger Chamberlin, 1910:244-245, pi. 36, figs. 1-5. Attems, 
1940:159. Chamberlin and Hoffman, 1958:73. 
Polydesmus amandus Chamberlin, 1910:249-250, pi. 38, fig. 6, pi. 
39, fig. 1. NEW SYNONYMY. 
Scytonotus orthodox Chamberlin, 1925:61. Attems, 1940:159. 
Chamberlin and Hoffman, 1958:73. Kevan, 1983:2969. NEW 
SYNONYMY. 
Archipolydesmus amandus: Attems, 1940:154-155. fig. 226. 
Scytonotus amandus: Chamberlin, 1943:143. Causey, 1954:223. 
Chamberlin and Hoffman, 1958:72. Loomis and Schmitt, 1971:117. 
Kevan, 1983:2969. 
Type specimens — One male and one female syntype (NMNH) 
taken by an unknown collector on an unknown date in the upper 
reaches of Mill Creek Canyon, Salt Lake County, Utah. 
