Genus Scytonotus 
21 
W. Ivie (NMNH), Ft. Douglas, F, 17 July 1951, Y. M. Wang 
(NMNH), and Mill Cr. Cyn., M, F, date and collector unknown 
(NMNH), and F, date unknown, R. V. Chamberlin (NMNH) TYPE 
LOCALITY; and Lamb Cyn., F, 20 September 1930, collector un- 
known (NMNH) and juvs., 21 July 1942, S. and D. Mulaik (NMNH). 
The following literature records are also deemed reliable and 
are designated by open symbols in figs. 9-10. 
MONTANA: Lincoln Co ., 8.5 mi (13.6 km) W Libby, and 
along Zulu Cr., S Fork Yaak R. (Loomis and Schmitt 1971). Min- 
eral Co ., Saltese (Loomis and Schmitt 1971). 
IDAHO: Bear Lake Co ., 7 mi (11.2 km) NW Georgetown 
(Chamberlin 1943). 
WYOMING: Teton Co., Moose (Causey 1954 a). 
Remarks — With two teeth each on the distal laminas, S. piger 
and columbianus are very similar. I have seen only three males of 
the latter, so my concepts may need to be modified when more 
specimens are available. From this material, the endomerite and 
tibiotarsus are consistently subequal in length, closely appressed to- 
gether, and overlap basally in S. piger, whereas in S. columbianus, 
the endomerite is much shorter and widely separated from the tibio- 
tarsus (Figs. 5-8). Additionally, the medial lamina is just a short 
flange that does not overhang the inner margin of the endomerite 
in S. columbianus, whereas it is much longer and extends into a 
basal lobe that does overhang this margin in S. piger. In nonsexual 
features, the tubercles are much stronger and more clearly demar- 
cated from each other in S. columbianus; they are smaller, more 
lowly rounded, and tend to merge with one another in S. piger. 
Similarly, the paranotal margins are much more deeply indented and 
emarginate in S. columbianus, in contrast to the nearly smooth con- 
dition in S. piger. 
Scytonotus columbianus Chamberlin 
Figs. 7-8 
Scytonotus columbianus Chamberlin, 1920:166-167, fig. 16. Attems, 
1940:158. Chamberlin and Hoffman, 1958:72. Kevan, 1983:2969. 
Shelley, 1990:20. 
Type specimens — Male holotype (MCZ) taken by J. B. Tyrrell, 
26 September 1883, in the Columbia Valley (probably the valley of 
the Columbia River), British Columbia, Canada. One male and one 
female paratypes (MCZ) taken by same collector, 1 September 1883, 
from a “swamp, tobacco plain” in the same locality. 
Diagnosis — Tibiae of legs 13-20/22 in males without distal 
lobes; endomerite much shorter than tibiotarsus, segregated from the 
