22 
Rowland M. Shelley 
latter for most of length; medial lamina a short, inconspicuous, 
broadly rounded flange at level of pulvillus, not overhanging inner 
margin of endomerite; distal lamina with widely separated, poorly 
demarcated, subacuminate teeth, proximal one larger; lateral 
lamina moderately broad, overhanging inner margin of endomerite 
(Figs. 7-8); females with paranota of segments 5-9 not reduced. 
Variation — The flange-like medial lamina is narrower and 
barely detectable on the male from Yoho National Park, and the 
proximal tooth on the distal lamina is much larger than the apical 
tooth in this individual. 
Ecology — The specimens that I collected in Mt. Revelstoke and 
Kootenay National Parks, British Columbia, were encountered under 
wet deciduous litter near streams. The male from Washington was 
recovered from a pitfall trap in an old field. 
Distribution — Extending from the Selkirk and Rocky Mountains 
at Revelstoke and Yoho National Parks, British Columbia, to the 
northeastern corner of Washington, ranging westward into the Co- 
lumbia River Valley (Fig. 10). Specimens were examined as fol- 
lows: 
CANADA: BRITISH COLUMBIA: Mt. Revelstoke Nat. Pk., 
Skunk Cabbage area, 2 juvs., 9 August 1989, R. M. Shelley (NCSM). 
Yoho Nat. Pk., 2 mi (3.2 km) S Takkakaw Falls, M, 5 October 
1963, A. Nimmo (VMNH) and Kicking Horse Camp, F, 5 October 
1963, D. R. Whitehead (VMNH). 10 mi (16 km) E Golden, F, 2 
juvs., 28 June 1988, S. & J. Peck (NCSM). Kootenay Nat. Pk., 
trail to Cobb Cpgd., juv., 7 August 1989, R. M. Shelley (NCSM). 
Columbia Valley, site not specified, 2M, F, 1 and 26 September 
1883, J. B. Tyrrell (MCZ) TYPE LOCALITY. Kaslo, F. 18 juvs., 
30 June-4 July 1903, A. M. Caudell (NMNH). 16 km E. Salmo, 
along Hwy. 3, ca. 1 mi (1.6 km) E jet. Hwy. 6, 2 juvs., 6 August 
1989, R. M. Shelley (NCSM). 
USA: WASHINGTON: Pend Oreille Co., ca. 7 mi (11.2 km) 
S Usk, Deer Cr., along WA hwy. 211 just S Davis L., 48.210°N, 
117.289°W, 6M, F, 29 August-9 September 1980, R. Crawford 
(UWBM). 
The Bergrothi Group 
Aside from the allopatric population of S. insulanus in eastern 
Washington, the bergrothi group is found exclusively along the Pa- 
cific Ocean. It is the only lineage occurring west of the Cascades, 
and S. bergrothi extends across the crest and onto the eastern slope 
of these mountains in Yakima and Kittitas counties, Washing- 
ton. Anatomically, the endomerites are longer than the tibiotarsi be- 
cause the distal lamina is prolonged to varying degrees. The latter 
