Genus Scytonotus 
23 
also bends sharply or curves so that it is angular with respect to 
the endomerite stem, and it is divided in S. insulanus. The medial 
laminas exhibit a distal tooth in S. bergrothi and simplex, and the 
lateral laminas are short with rounded lobes except on S. simplex, 
which possesses a lightly serrate basal lobe and a distal tooth. 
Scytonotus bergrothi and insulanus are broadly sympatric in coast- 
al Washington, northwestern Oregon, and southwestern British 
Columbia, whereas S. simplex is parapatric to the south, with only 
minimal overlap of S. bergrothi (Figs. 20, 32). I am unable to 
resolve the relationships among the components and show them in 
Figure 34 as an unresolved trichotomy. 
Components — bergrothi Chamberlin, insulanus Attems, simplex 
Chamberlin. 
Scytonotus insulanus Attems 
Figs. 11-12 
Scytonotus insulanus Attems, 1931:147-149, figs. 240-245; 1940:157- 
158, figs. 229-231. Chamberlin and Hoffman, 1958:73. Kevan, 
1983:2969. 
Scytonotus insulans : Shelley, 1990:20. 
Type specimens — One male and one female syntypes (NMV) 
taken by an unknown collector on an unspecified date in 1934 at 
Nanaimo, Vancouver Island, British Columbia, Canada. 
Diagnosis — Tibiae of legs 13-22 in males with distal lobes; 
endomerite much longer than tibiotarsus; medial lamina relatively 
short, with moderately acuminate flange overhanging inner margin 
of endomerite; distal lamina apically divided, medial branch slightly 
expanded and broadly rounded basally, tapering into greatly pro- 
longed, strongly decurved projection, tip narrowly rounded, lateral 
branch about 1/3 as long as latter, expanding distad and tapering 
slightly to subacuminate tip; lateral lamina expanding into moderate- 
size, narrowly rounded lobe, overhanging inner margin of endomerite 
(Figs. 11-12); paranota of segments 5-9 reduced in females. 
Variation — There is very little variation among the gonopods 
of S. insulanus. In the male from 12 mi (19.2 km) east of Salem, 
Marion County, Oregon, the flange on the medial lamina is nar- 
rower and more rounded than that of the type, and the lobe on the 
lateral lamina is smaller, more pointed, and located more proximad. 
A few other individuals exhibit similar, minor variation in the size 
of these structures, but most gonopods agree closely with the type. 
Ecology — Labels with preserved samples give the following 
microhabitat information for S. insulanus : alder and birch litter, 
