26 
Rowland M. Shelley 
USA: ALASKA: Yakutat Bay (Cook 1904). 
WASHINGTON: Clallam Co., Elwha (Causey 19546). 
Remarks — The records from the Columbia Plateau Physiographic 
Province in Whitman County, Washington, are quite surprising, but 
the data appear accurate, and the adults conform to all the diagnos- 
tic traits of S. insulanus. This is the only record of the genus from 
southeastern Washington, and it suggests occurrence of S. insulanus 
in the Hell’s Canyon area of northeastern Oregon. The Lane and 
Douglas county Oregon sites, about 90 mi (144 km), from the clos- 
est locality, in Benton County, also may represent an allopatric 
population. 
Scytonotus bergrothi Chamberlin 
Figs. 13-14 
Scytonotus sp. Cook, 1904:pl. 4, figs. 2a-d. 
Scytonotus bergrothi Chamberlin, 1911:262-264, fig. 16. Attems, 
1940:158. Chamberlin and Hoffman, 1958:72. Kevan, 1983:2969. 
Scytonotus pallidus Attems, 1931:145-147, figs. 234-239; 1940:156- 
157, figs. 227-228. Causey, 1954:82. Shelley, 1990:20. 
Type specimens — One male and 3 female syntypes (NMNH) 
collected by D. E. Bergroth on an unknown date at Bremerton, 
Kitsap County, Washington. 
Diagnosis — Tibiae of legs 13-21 in males with distal lobes; 
endomerite slightly longer than tibiotarsus; medial lamina expand- 
ing into broad flange for most of length, terminating in sharply 
acute distal tooth; distal lamina with smaller, sharply acute tooth 
proximad, narrowing, curving gently, and slightly prolonged thereaf- 
ter, apically acuminate, with or without variable undulations, short 
teeth, or spurs on outer margin; lateral lamina relatively short, ex- 
panding into moderately large, broadly rounded lobe, overhanging 
inner margin of endomerite (Figs. 13-14); paranota of segments 5-9 
reduced in females. 
Variation — The most notable gonopodal variation are the en- 
largements of the undulations on the outer margin of the distal 
lamina in males from British Columbia. Barely noticeable in south- 
ern specimens, the undulations become larger and more distinct around 
Vancouver and on Vancouver Island, and even are denticulate on a 
few males. Other gonopodal variation involves minor differences in 
the length and position of the spur on the inner margin of the 
endomerite near the hairpad. It projects well beyond the hairpad 
and is visible in lateral view in some males but is shorter in oth- 
ers; it can also be distal or proximal to the pulvillus. The relative 
sizes of the distal tooth on the medial lamina and the basal tooth 
