Genus Scytonotus 
31 
Scytonotus simplex Chamberlin 
Figs. 15-16 
Scytonotus simplex Chamberlin, 1941:16, pi. 3, fig. 30. Chamberlin 
and Hoffman, 1958:73. Kevan, 1983:2969. 
Type specimens — Male holotype, female allotype, and one male 
and one female paratypes (NMNH) collected by J. C. Chamberlin, 
18 November 1939, at Days Creek, Douglas County, Oregon. 
Diagnosis — Tibiae of legs 13-20 in males with distal lobes; 
endomerite moderately longer than tibiotarsus; medial lamina expand- 
ing into moderate-size flange for proximal 2/3 of length, indenting 
broadly, then expanding into moderate-size, acuminate, distal tooth; 
distal lamina bent nearly perpendicular to tibiotarsus, narrowly ex- 
panded basally, inner margin curving slightly bisinuately into sub- 
acuminate tip; lateral lamina located well distal to pulvillus, expanding 
into moderate-size basal lobe with lightly irregular margin, narrow- 
ing slightly, then expanding into sharply acute tooth distal to mid- 
length (Figs. 15-16); paranota of segments 5-9 reduced in females. 
Variation — The distal lamina and the distal parts of the medial 
and lateral laminas are closely similar on all specimens, but the 
proximal parts of the last two lamellae vary considerably. The 
breadth of the basal flange on the medial lamina is often narrow- 
er than in the illustrated male, and an individual from Del Norte 
County, California, has a sharply acute tooth at the corner of the 
flange. The distal tooth on the medial lamina is also much larger 
in the male from San Joaquin County, California. On the lateral 
lamina, the basal serrated lobe varies considerably. In a male from 
Coos County, Oregon, it is longer and located more proximad; it is 
also longer and narrower in individuals from Trinity County, Cali- 
fornia. Conversely, it is much shorter and narrower in the San Joaquin 
County male and is barely detectable in one from Del Norte County. 
Ecology — Labels with preserved samples indicate the following 
microhabitats for S. simplex : pine tree trunks after rain, hemlock 
duff, birch litter, and under wet leaves. 
Distribution — Along the Pacific Coast from northern Lincoln 
County, Oregon, to San Francisco Bay and the northern San Joaquin 
Valley, California (Fig. 20). The range extends eastward onto the 
western slope of the Cascades in Lane, Douglas, and Jackson coun- 
ties, Oregon, and overlaps slightly both the main area and the Lane 
County population of S. insulanus (Fig. 31). Specimens were exam- 
ined as follows: 
OREGON: Lincoln Co., 0.6 mi (0.9 km) NW Elk City, along 
Yaquina R., M, F, 20 December 1971, E. M. Benedict (WAS). 5.5 
mi (8.8 km) E, 2.5 mi (4.0 km) S, Tidewater, along Alsea R., 3M, 
