Genus Scytonotus 
35 
km) W Inverness, 2F, 1 May 1976, J. T. Doyen (CIS); and Inverness 
Ridge, M, 17 November 1962, N. B. Causey (FSCA) and MM, FF, 
9 October 1963, J. S. Bucket (UCD, FSCA). San Joaquin Co., 
Stockton, M, 27 October 1973, L. S. Hawkins (NCSM). 
Remarks — The name of this species is puzzling and clearly a 
misnomer; the gonopod of S. simplex is hardly “simple” either alone 
or in comparison with those of the other species that had been 
proposed by 1941. One wonders about Chamberlin’s reasoning, be- 
cause the only species with truly “simple” gonopods is S. inornatus. 
The Inornatus Group 
A monobasic lineage, this group is characterized by an ab- 
sence of modifications that is most noticeable on the distal lamina. 
The medial lamina expands into a broad flange; the lateral lamina 
is long and slender; and the endomerite and tibiotarsus are subequal 
in length. The lineage is thus the most plesiomorphic in Scytonotus, 
and from its apparently restricted distribution, the lone component 
species appears to be a relict and the sole surviving remnant of its 
line. Such a nondescript, unmodified ancestral form could have given 
rise to both other lineages, and its occurrence on the eastern slope 
of the Cascades points to this general region as the likely source 
for the other lines and the primary center of evolution within Scytonotus. 
Component, inornatus, new species. 
Scytonotus inornatus, new species 
Figs. 17-18 
Type specimens — Male holotype and one female paratype (FSCA) 
collected by J. Schuh, 25 October 1972, along a canal at Geary 
Ranch (exact location unknown), Klamath County, Oregon; four fe- 
male paratypes (FSCA) taken by same collector at same locality on 
28 October 1971. 
Diagnosis — Tibiae of legs 13-20 in males with distal lobes; 
endomerite subequal in length to that of tibiotarsus; medial lamina 
expanding into large, broadly rounded flange for most of length, 
overhanging and extending well beyond inner margin of endomerite; 
distal lamina with inner margin smoothly linear, without denticula- 
tions or other modifications, outer margin narrowing abruptly to 
subacuminate termination with inner; lateral lamina long and nar- 
row, expanding slightly proximad and distad but not overhanging 
inner margin of endomerite; paranota of segments 5-9 of females 
reduced. 
Description — Head normal for genus, densely covered with short, 
fine, parallel-sided setae; epicranial suture distinct; genae extending 
well beyond adjacent cranial margins. 
