Genus Scytonotus 
37 
on the lateral lamella, S. inornatus is the most pleisiomorphic known 
species. A hypothetical ancestral Scytonotus would resemble S. inornatus 
with a reduced medial lamina, and all other species of Scytonotus 
could have arisen from such an ancestral form. The species occurs 
east of the crest of the Cascade Mountains in the foothills on the 
boundary with the arid Basin and Range Physiographic Province. 
This area is much drier than those west of the crest where S. 
simplex and bergrothi occur. Presumably, the Geary Ranch is near 
Klamath Falls and is therefore in the Klamath River drainage, which 
flows westward and empties into the Pacific Ocean in southern Del 
Norte County, California. 
Scytonotus sp. 
To completely detail the generic distribution in the West, I 
record here the samples of Scytonotus that lack adult males and 
cannot be assigned to a species on a geographic basis. In Canada, 
coastal Washington, and Oregon, the records could refer to either 
S. insulanus or bergrothi, and the Stevens County, Washington, 
literature record (Causey 1954b) could be either S. piger or S. 
columbianus. These records are indicated by question marks in 
Figures 10 and 19-20. 
CANADA: BRITISH COLUMBIA: VANCOUVER ISLAND: 
Courtenay, F, juv., June 1965, N. L. H. Kraus (NMNH). Robson 
Bight, Tsitika R., 2 juvs., 28 July 1986, D. H. & J. L. Kavanaugh 
(CAS). Little Qualicum Falls Prov. Pk., 2 juvs., 30 July 1989, R. 
M. Shelley (NCSM). 5 km NE Port Renfrew, 2F, 31 July 1989, R. 
M. Shelley (NCSM). Goldstream Prov. Pk., F, 2 August 1989, R. 
M. Shelley (NCSM). 
MAINLAND SITES: Shannon Falls Prov. Pk., FF, juvs., 29 
July 1989, R. M. Shelley, (NCSM). Along hwy. 99, 132 mi (21 
km) N Squamish, FF, juvs., 29 July 1989, R. M. Shelley (NCSM). 
Capilano Cyn. Reg. Pk., 4F, 28 July 1989, R. M. Shelley (NCSM). 
1.9 mi (3 km) SE Elope, along Silver Skagit Rd., F, 30 June 1988, 
S. & J. Peck (NCSM). Ten Mile, exact location unknown, juv., 20 
June 1903, R. F. Currie (NMNH). 
USA: WASHINGTON: Jefferson Co., Olympic Nat. Pk., Graves 
Cr. Cpgd., 2F, 25 August 1990, R. M. Shelley (NCSM). Grays 
Harbor Co., along Moclips R., 6.4 mi (10.2 km) W US hwy. 101, 
F, juv., 25 August 1990, R. M. Shelley (NCSM). 
OREGON: Hood River Co., Perham Cr., 5F, 2 juvs., 4 Au- 
gust 1929, collector unknown (NMNH); and 4 mi (6.4 km) S Parkdale, 
