40 
Rowland M. Shelley 
Type specimens — Not known to exist, although part of the type 
series may be extant at the BMNH. The type locality is the vicin- 
ity of Philadelphia, Philadelphia County, Pennsylvania. 
Diagnosis — Tibiae of legs 13-20 in males with distal lobes; 
tibiotarsus and endomerite essentially subequal in length, former slightly 
longer; medial lamina expanded into broadly rounded flange extend- 
ing most of length of lamina, overhanging inner margin of endomerite; 
distal lamina with minute apical tooth preceded by two narrowly 
segregated teeth; lateral lamina expanding into narrow, flange, mar- 
gin irregular, with one or two variable teeth (Figs. 21-22); females 
with paranota of segments 5-9 reduced. 
Variation — Hoffman (1962b) noted that S. granulatus maintains 
its structural integrity over the entire range and that he could not 
discern geographically variable details. In spot-checking samples from 
across the range, I can corroborate Hoffman’s observation: I found 
only minor differences in the sizes of the flanges on the medial 
and lateral laminas as well as in the size and degree of separation 
of the teeth on the distal lamina. 
Ecology — I have encountered S. granulatus in a variety of mi- 
crohabitats and believe it can be expected in most moist environ- 
ments within its range; those with a preponderance of deciduous 
litter are more probable than those with a higher proportion of 
pine litter. The milliped will likely be found well within the litter, 
on the undersides of leaves or pieces of wood and bark. 
Distribution — Covering almost the entire generic distribution in 
eastern North America, ranging from the vicinities of Trois Rivieres, 
Quebec, and Sault St. Marie, Ontario, to central South Carolina, 
central Tennessee, and northeastern Arkansas, and east/west, from 
Vermont, coastal Virginia, and the Outer Banks of North Carolina 
to eastern Kansas and Nebraska (Fig. 29). The only part of the 
eastern generic range that is not inhabited by S. granulatus is the 
Blue Ridge Province from northern Virginia to north Georgia, where 
it is replaced by the endemic species, S. virginicus and australis. 
The distribution encompasses parts or all of a dozen physiographic 
provinces and 16 states, 6 of which — Pennsylvania, West Virginia, 
Kentucky, Ohio, Indiana, and Illinois — are wholly within its range. 
Specimens were examined as summarized below. Complete locality 
data are provided for states and Canadian provinces where S. 
granulatus is known from five or fewer counties; the latter are 
listed in alphabetical order when the milliped has been collected in 
more than five counties. 
CANADA: QUEBEC: Drummond Co., Drummondville, MM, FF, 
