50 
Rowland M. Shelley 
TENNESSEE: Sevier Co., Gatlinburg, M, F, date and collector 
unknown (NMNH). 
SOUTH CAROLINA: Oconee Co., Clemson, F, 22 December 
1966, J. A. Payne (VMNH). Pickens Co., Burnt Mtn., F, 6 Novem- 
ber 1960, L. Hubricht (VMNH). 
GEORGIA: White Co., 2.7 mi (4.3 km) N Robertsville, F, 12 
March 1961, L. Hubricht (VMNH). Dawson Co., 6 mi (9.6 km) W 
Amicalola Falls, 3M, 8F, 6 November 1960, L. Hubricht (NMNH, 
VMNH) TYPE LOCALITY; and 6 mi (9.6 km) SW ' Dawsonville, 
F, 8 April 1961, L. Hubricht (VMNH). 
Remarks — The record of juveniles from Highlands, Macon 
County, North Carolina, that Hoffman (1950a, 19626) assigned to 
S. granulatus and michauxi, respectively, is properly referrable to S. 
australis. 
Scytonotus australis X S. virginicus michauxi hybrid 
The gonopods of a male from the Great Smoky Mountains 
National Park, North Carolina/Tennessee, show features of both S. 
australis and v. michauxi. The distal lamina is clearly that of the 
former; the lateral lamina is clearly that of the latter; and the me- 
dial lamina combines aspects of both, with a distal tooth like that 
of the population of S. australis in the “Smokies,” and with a 
smaller, proximal tooth so that the lamella has two teeth as in S. 
v. michauxi. As all other samples from this area are clearly one 
species or the other, I do not think this one individual suffices to 
overturn my conclusion that these forms are specifically distinct; 
hence, I record it as a hybrid. The sample emphasizes the need for 
more material from the borderline counties in southwestern North 
Carolina and southeastern Tennessee. Data are as follows: the site 
is denoted by a question mark in Figure 30. 
NORTH CAROLINA/TENNESSEE: Swain/ Sevier cos., Clingman’s 
Dome summit, M, F, 24 October 1969, W. A. Shear (WAS). 
Distribution 
The generic range is characterized earlier in this work and 
need not be repeated, but it is informative to examine the distribu- 
tions of the species and lineages (Figs. 31-33), and their spatial 
relationships. As shown in Figures 31-32, the species-group taxa 
demonstrate allopatric and parapatric spatial relationships with mini- 
mal range overlaps, except for the high level of sympatry between 
S. insulanus and bergrothi (Fig. 31, nos. 1 and 2, respectively) on 
Vancouver Island, the southwestern mainland of British Columbia, 
and western Washington. Large gaps exist between S. granulatus 
in the east and S. piger in the west (about 984 mi [1590 km]), 
