Genus Scytonotus 
51 
Fig. 32. Comparative distributions of species and subspecies of Scytonotus 
in eastern North America. 7, granulatus\ 8, australis ; 9, v. michauxi; 10, 
v. virginicus\ 11, virginicus intergrades. 
between the two areas occupied by S. piger (approximately 274 mi 
[438 km]), and between the main range and the allopatric popula- 
tion of S. insulanus in southeastern Washington (around 208 mi 
[333 km]). Smaller distances separate apparently allopatric popula- 
tions of S. insulanus and bergrothi in Oregon’s Lane and Douglas 
counties, respectively, from the main areas of their ranges. Scytonotus 
is therefore cohesive along the Pacific Coast and demonstrates at- 
tributes of a small mosaic (see Shelley and Whitehead 1986, Shelley 
