Genus Scytonotus 
53 
Fig. 33. Comparative distributions of species groups of Scytonotus. 1, the 
inornatus group; 2, the bergrothi group; 3, the granulatus group. The 
boundary line of the bergrothi group is continued through the ocean off 
British Columbia and Alaska to show that the range excludes the Queen 
Charlotte Islands. 
did not formally propose, separate family status. Hoffman (19626) 
mentioned that this two-branched development might be homologous 
to that in some European species of Polydesmus, for example P. 
inconstans Latzel, which has been introduced into North America 
and occurs in many American and Canadian cities. Golovatch (1991) 
suggested that the Asian genera Schizoturanius Verhoeff and 
Turanodesmus Lohmander, from Kirgizstan and Turkestan, respec- 
tively, might be allied to Scytonotus because they also possess the 
two-branched structure. As illustrated by Attems (1940), the gonopods 
of T. stummeri (Attems) and T. inermis Lohmander resemble the 
basic structure of Scytonotus , so Golovatch’s idea seems plausible 
