New Crayfish Species 
69 
of merus with longitudinal row of 2 to 8 (mode 4-5) prominent, acute 
spines in addition to long distal spine, ventrolateral ridge with longitudinal 
row of 2 to 7 (mode 2-3) prominent, acute spines in addition to long 
distal spine. Ventral surface of carpus of cheliped almost always with 
strong distomedian spine, and larger acute spine on lateral articular 
condyle; mesial surface of carpus with long, procurved spine at about 
midlength, usually smaller spine distal to it, and tubercle, often spiniform, 
near proximal margin. 
Hook on ischium of third pereiopod of form I male usually extending 
beyond basioischial articulation, sometimes opposed by low, vestigial 
tubercle on basis. Length of gonopod (first pleopod) of form I male 
divisible into TCL 2.1 to 2.5 ( * = 2.2, n - 54) times; cephalic surface 
of gonopod with prominent angular shoulder, distal to which both 
rami inclined cephalically; terminal elements long, slender, subparallel 
to about midlength of mesial process; central projection tapering smoothly 
from base to acute tip, distal portion gently recurved, tip directed 
caudodistally and reaching to or beyond midlength of coxa of first 
pereiopod when abdomen flexed; length of central projection 46.5 
to 57.9 ( x = 51.6, n = 54) percent of total length of gonopod; tip 
of mesial process somewhat spatulate and mildly excavate cephalically; 
length of mesial process 34.7 to 44.5 (* = 41.5, n = 54) percent of 
total length of gonopod, and 73.3 to 89.2 (x = 80.4, n = 54) percent 
of length of central projection. In caudal (ventral) aspect, bases of 
gonopods and tips of mesial processes contiguous; tip of central projection 
directed slightly distolaterally, that of mesial process directed distomesially. 
Annulus ventralis inflexibly fused to sternum; annulus 1.3 to 
1.4 times wider than long, with deep, broad transverse fossa; cephalic 
border of annulus convex, caudal border subangular; cephalomedian 
wall with short longitudinal trough, flanked at each side by ridge, 
ridges diverging caudally and each terminating in large, rounded cephalo- 
lateral prominence, prominences in ventral aspect obscuring cephalic 
half or more of fossa; caudomedian wall dissected by sinus; obvious 
sulcus cephalic to caudal margins. 
Measurements of type specimens provided in Table 1. 
Holotypic male, form I — Body and eyes pigmented, eye 2.5 mm 
diam. Cephalothorax (Fig. 1A, D) subcylindrical, somewhat depressed 
dorsally; carapace widest at midlength, width greater than depth at 
caudodorsal margin of cervical groove; abdomen narrower than carapace. 
Areola 4.1 times longer than wide, constituting 29.5 percent of TCL 
(42.0% of PCL), with 5 punctations across narrowest part and low, 
rounded eminence at caudal base. Rostrum acarinate, length 31.2 percent 
of TCL (44.4% of PCL), deeply excavate, with thickened, elevated 
