72 
John E. Cooper and Martha Riser Cooper 
Postorbital ridge strong throughout most of length, with shallow, 
setiferous dorsolateral groove; cephalic margin terminating in strong 
spine. Suborbital angle nearly obsolete, delimited ventrally by shallow 
rounded notch at proximal base of antennal peduncle. Cervical spines 
strong, acute, directed cephalolaterally, 1 each side, plus single, small 
spiniform tubercle just dorsal to spine. Branchiostegal spine prominent, 
acute. Hepatic region punctate, with some small subspiniform tubercles. 
Carapace densely punctate dorsally (including gastric region) and laterally, 
somewhat granulate ventrolaterally; cluster of tubercles just caudal 
to branchiostegal spine and ventral to cephalolateral portion of cervical 
groove, which with row of tubercles extending along ventral margin; 
cervical groove deep, uninterrupted, with shallow, curved sulcus (tributary 
to groove) ventral to cervical spine, creating subglabrous mound below 
spine. 
Abdomen slightly longer than carapace; pleura well developed, 
subtruncate, with smoothly rounded cephaloventral margins and sub- 
angular caudoventral margins; minute atypical notch in apex of second 
through fourth pleura. Cephalic section of telson with large, acute, 
immovable spine at caudolateral corner, and smaller, movable spine 
just mesial to it (that on right side small, regenerate); cephalic and 
caudal sections of telson partly separated by oblique lateral incisions 
and shallow transverse sulcus. Promixal podomere of uropod with long, 
corneous spine on mesial lobe and slightly smaller spine on lateral 
lobe; mesial ramus of uropod with distolateral marginal spine and 
relatively broad median keel terminating distally in acute premarginal 
spine; cephalic section of lateral ramus with median keel terminating 
in acute spine at transverse flexure, which with total of 13 fixed spines 
and 1 large movable spine at lateral margin. 
Cephalic lobe of epistome (Fig. 1H) spatulate, tilted cephaloventrally, 
with moderate constriction and deep transverse groove at caudal base; 
cephalomedian margin with slight notch ventral to short projection; 
cephalolateral margins elevated (ventrally), broad, sloping onto ventral 
surface and with thin, emarginate rim bearing short setae; lateralmost 
extremities mildly flanged; ventral surface of lobe concave, punctate, 
with short, sparse setae, concavity continuing into cephalomedian notch; 
caudal one-third of ventral surface with low, subtriangular eminence; 
main body of epistome relatively glabrous, cephalomedian and cephalo- 
lateral margins forming a somewhat hemitubular, curved ridge; central 
depression of body broad, deep, with deep fovea situated in midline 
at cephalic margin of depression; zygoma moderately arched, wider 
than space between renal apertures; pits at cephalolateral borders of 
zygoma elongate and relatively shallow. 
