110 
Rowland M. Shelley 
Jeekel, 1971:41. Hoffman, 1980:73. 
Hypozonium Cook, 1904:62. Cook and Loomis, 1928:17. Chamberlin 
and Hoffman, 1958:187. Buckett, 1964:29. Jeekel, 1971:39. 
Hoffman, 1980:73. Kevan, 1983:2962. NEW SYNONYMY. 
Euzonium Chamberlin, 1950:1. Chamberlin and Hoffman, 1958:187. 
Buckett, 1964:29. Jeekel, 1971:38. Hoffman, 1980:73. NEW 
SYNONYMY. 
Type species — Of Octoglena, O. bivirgata Wood, 1864, by monotypy; 
of Hypozonium , H. anurum Cook, by monotypy; of Euzonium , E. crucis 
Chamberlin, 1950, by original designation. 
Diagnosis — Dorsum granular, with or without three dark, longitudinal 
stripes; caudal metatergal margins detached from succeeding tergite, 
upturned to varying degrees, body broad (W/L ratio generally 28- 
38%), flattened “bell shaped” in profile, sides extending strongly laterad; 
collum broad, overhanging epicranium and at least one pair of ocelli; 
telson variable, either broad and comprising nearly entire caudal width, 
or relatively narrow and comprising around half of caudal width; head 
subtriangular, with three ocelli on each side arranged linearly in angular 
rows beginning at levels of antennal sockets; sternum of anterior gonopods 
with strong, apically hirsute lobes, segregated to varying degrees; anterior 
gonopods curving variably anteromediad distad, ultimate podomere 
divided, with broad, hirsute, ventral lobe of variable size usually over- 
hanging dorsal glabrous branch, latter either short, broad, and apically 
blunt, curved slightly ventrad, and directed anteromediad, or long, 
narrow, and acuminate, slightly sinuate and directed sublaterad; corners 
of 4th and 5th podomeres extended on caudal side; coxa with or without 
hirsute anterior lobe, length variable; posterior gonopod with ultimate 
podomere simple and acicular, apically acuminate, fimbriate, or lightly 
hirsute, projecting anteriad between solenomere and ventral lobe. 
Species — Five. 
Distribution — Along the Pacific Coast from the southwestern corner 
of the British Columbia mainland to central Santa Cruz County, California, 
extending inland to the western slope of the Cascade Mountains from 
British Columbia to central Oregon and the eastern slope of the Coast 
Range in southern Oregon and California, with a localized allopatric 
species some 75 mi (120 km) to the east in the Sierra Nevada foothills, 
Placer County, and one 1,897 mi (3,035 km) farther east in the eastern 
United States, extending from westcentral South Carolina to southcentral 
Tennessee and northwestern Alabama (Figs. 1, 28-29). The coastal 
species are contiguous and demonstrate parapatric spatial relationships 
and a sublinear, north to south, arrangement, the area being wider 
from central Oregon northward. Dimensions are approximately 850 
