Hirudisomatid Millipeds 
111 
mi (1,360 km), north/south, and 110 mi (176 km), east/west, for the 
coastal area, and 220 mi (352 km), north/south, and 380 mi (608 
km), east/west, for the eastern. 
Relationships — With the somatic differences and the apomorphic 
absence of the coxal lobe, O. gracilipes is sister to the western species 
(Fig. 30). Octoglena sierra is sister to those along the Pacific Coast, 
which share the configuration of the ultimate anterior gonopod podomere, 
and O. anura, with the short, apically linear coxal lobe, is sister to 
0. prolata plus O. bivirgata, in which the lobe is extended and apically 
rounded. 
Remarks — The anterior gonopods of each species are quite uniform 
and show little intraspecific variation, so I analyze segment numbers 
and lengths of measurable specimens in the variation sections. As 
per the recommendation of Enghoff et al. (1993), I exclude the telson 
from the segment counts. 
Key to Species of Octoglena 
1. Caudolateral corners of midbody metatergites slightly but distinctly 
extended and rounded; telson broad, extending for nearly entire 
breadth of caudal extremity; anterior gonopod coxae with hirsute 
lobes of varying lengths (Figs. 9, 13, 16, 19); southwestern 
British Columbia to central California 2 
Caudolateral corners of midbody metatergites not extended, 
continuous with middorsal margins, apically blunt; telson narrow, 
comprising no more than half of breadth of caudal extremity; 
anterior gonopod coxae without lobes (Fig. 25); North and 
South Carolina to Tennessee and Alabama gracilipes (Loomis) 
2. Ultimate podomere of anterior gonopod with broad, distinct lobe 
ventral to and overhanging dorsal glabrous branch; latter rela- 
tively short and broad, curved slightly ventrad, directed antero- 
mediad (Figs. 9, 13, 16); along Pacific Coast from British 
Columbia to Santa Cruz County, California 3 
Ultimate podomere of anterior gonopod with at most short, 
indistinct, ventral lobe, not overhanging dorsal branch; latter 
relatively long and narrow, slightly sinuate, directed sublaterad 
(Fig. 19); western Placer County, California 
sierra , new species 
3. Coxal lobe of anterior gonopod short and apically sublinear, not 
overlying distal podomeres; ultimate podomere of posterior 
gonopod apically narrow and with numerous short hairs (Figs. 
13, 15); southwestern British Columbia to northern Douglas 
County, Oregon anura (Cook) 
