Hirudisomatid Millipeds 
129 
34 to 49 segments and vary in length from 7.1 to 11.7 mm, the shortest 
individual having 36 segments and being 0.7 mm shorter than that 
with the least segments. Female segment numbers vary from 38 to 
43 and lengths, from 9.8 to 11.9, the latter, of an individual with 
41 segments, being 0.4 mm longer than the female with the most 
segments (table 4). 
Ecology — One paratype sample was encountered in “litter under 
oak.” 
Distribution — A localized species known only from the type and 
paratype localities in the foothills of the Sierra Nevada (Fig. 28), 
O. sierra is detached from the coastal representatives and occurs some 
75 mi (120 km) east of the nearest site of O. bivirgata, in Colusa 
County. It is the easternmost western species and occupies an inter- 
mediate geographical position, albeit far to the western side of the 
generic range. By combining gonopodal attributes of O. gracilipes, 
the long, sinuate dorsal branch and the indistinct ventral lobe, with 
a trait of the coastal species, the coxal lobe, O. sierra links the ana- 
tomical extremes, which justifies congeneric status. 
Octoglena gracilipes (Loomis), new combination 
Figs. 22-27 
Polyzonium gracilipes Loomis, 1971:157-159, figs. 18-23. 
Type specimens — Male holotype and one male and one female 
paratypes (NMNH) and one male and one female paratypes (FSCA) 
collected by H. R. Steeves, 17 June 1962, at Cloudland Canyon State 
Park, Dade County, Georgia. The NMNH sample also includes one 
male and three female polyzoniids. 
Diagnosis — Dorsum without stripes, color pale yellow to white; 
caudal metatergal margins moderately upturned, caudolateral corners 
of midbody metatergites not extended, contiguous with middorsal margins, 
blunt; telson narrow, comprising about half of breadth of caudal extrem- 
ity; sternal lobes of anterior gonopods relatively long, narrowly segregated; 
coxae of latter without lobes; dorsal branch of ultimate podomere 
long, narrow, and sinuate, apically acuminate, directed sublaterad; ventral 
lobe of ultimate podomere indistinct, not overhanging dorsal branch; 
ultimate podomere of posterior gonopod apically narrow and attenuated 
(Figs. 22-27). 
Variation — Measurable males with seemingly mature gonopods 
have from 19 to 33 segments and vary in length from 2.8 to 7.9 
mm; female segment numbers vary from 13 to 35, while lengths vary 
from 1.7 to 8.5 mm. These data are presented in table 5 with localities 
arranged in a general east to west sequence. Lengths appear roughly 
