New Milliped Genus 
39 
The De La Howe woods is the “progenitor” of all piedmont 
forests. The canopy is dominated by loblolly pines on the slopes 
and in the draws with shortleaf pine on the ridges with several age 
classes and trees 12-42” D. B. H. The subcanopy is a mosaic of 
oaks and cover hardwood elements . . . On the ridges and slopes 
are red, white, black, post and southern red oaks along with 
pignut and mockernut hickories, red mulberry, yellow poplar and 
sweet gum; in the draws and sheltered slopes are beeches and 
southern sugar maple. Most of the subcanopy trees vary from 12- 
24” D. B. H. Beneath the subcanopy is a layer of hop hornbeam, 
storax and flowering dogwood along with some tall blue haws 
( Viburnum prunifolium). The shrub layer is composed of 
leatherwood in the draws, dwarf pawpaw, aromatic sumac, and 
maple-leaved viburnum on the slopes and deerberry ( Vaccinium 
stamineum ) and New Jersey tea along the ridge. . . . the herb layer 
is scant with the woody layers above but it is a unique combina- 
tion of partridge berry, wintergreen and pennywort ( Obolaria 
virginica). 
This is a classic example of the eastern North American Oak- 
Pine Forest with a maximum woody plant diversity. Tree size for 
the age of the stand is not overly impressive but the combination 
of woody species over this nearly circumneutral soil derived from 
a basic igneous rock and the abundance of the Obolaria make this 
a truly outstanding site. 
The tract is now recognized as a Natural Area by the Society 
of American Foresters. 
Distribution. — Brevigonus shelfordi ranges about 112 km (70 mi.) 
along the north side of the Savannah River in piedmont South Carolina, 
from the vicinity of Clemson to McCormick. It occurs about 32 km (20 
mi.) from the river into South Carolina, but most individuals have been 
collected less than 16 km (10 mi.) from the river in Abbeville and McCor- 
mick counties. The species has not been encountered in Anderson 
County and does not appear to occur in Georgia. I have collected along 
the Savannah River in Hart, Elbert, Lincoln, and Columbia counties, the 
Georgia counties bordering the range of shelfordi, without finding the 
milliped. The dominant xystodesmid in this area is Cleptoria abbotti 
Hoffman, which seems to replace shelfordi south of the Savannah River. 
In July 1979 an assistant and I collected shelfordi along South Carolina 
highway 72 in Abbeville County close to the Savannah River, but just 
across the river in essentially the same habitat in Elbert County, Georgia, 
we found only C. abbotti. I thus feel that shelfordi is absent from Georgia 
even though this area has not been sampled as thoroughly as have areas 
in South Carolina. 
Specimens have been examined from the following South Carolina 
localities. Unless otherwise indicated, all were collected by the author 
and deposited in the N. C. State Museum collection, the invertebrate 
catalogue number of which is shown in parentheses. Materials from the 
