New Beetle Species Groups 
91 
Pseudanophthalmus cumberlandus Valentine 
Valentine 1937:96. Type locality, Piper Cave, Smith Co., TN. 
The species is seasonally abundant in damp areas of Piper Cave, es- 
pecially on previously flooded muddy floors and around rimstone pools 
and bat guano, but it is usually rare or absent in winter. Unlike the ma- 
jority of the species in the group, P. cumberlandus is unusually robust and 
convex, with prominent humeri and nearly perpendicular prehumeral 
borders, the elytral striae very shallow. I have also taken it in Skeleton 
Cave, Smith County, and Ann White Cave, Macon County, Tennessee. 
Pseudanophthalmus productus, new species 
Fig. 5 
Etymology.— Latin productus, “produced.” 
Diagnosis. — Closely similar in the robust, convex body form and 
nearly perpendicular prehumeral borders to P. cumberlandus Valentine, 
differing in denser pubescence, wider pronotum base, deeper elytral 
striae, and long, produced, aedeagal apex. 
Description. — Length 4.2-5.4, mean 4.9 ± S.E. 0.3 mm (N =35). 
Form robust and convex, pubescent, rufotestaceous. Head about 0.1 
longer than wide; labrum doubly emarginate; antenna 0.7 body length. 
Pronotum 0.87 as long as wide, transverse-subquadrate; base wider than 
apex and 0.8 maximum width, which occurs in apical third; sides sinuate 
in basal sixth, but very shallowly so, hind angles about right, sometimes 
slightly acute or slightly obtuse, feebly emarginate behind, secondary 
angles present but rather subdued. Elytra 1.6 times longer than wide, 
prehumeral borders nearly perpendicular to midline, humeri prominent 
and serrulate, inner 6 striae moderately impressed, intervals subconvex, 
apical groove short and very rounded, joining third stria at level of an- 
terior apical puncture. Aedeagus 0.62-0.77, mean 0.70 ± S.E. 0.04 mm 
long (N = 30), moderately and evenly arcuate, apex elongate, slender, 
and much produced; copulatory sclerite elongate. 
Type series. — Holotype male (American Museum of Natural 
History) and 27 paratypes, Neil Fisher Cave (=Rip Van Winkle Cave), 
Smith Co., Tennessee, 25 March 1961, T. C. Barr; 33 paratypes, same 
cave, 18 October 1948, J. M. Valentine and W. B. Jones. 
Measurements (in mm).— Holotype, total length 4.80, head 0.92 long 
