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PSYCHE 
VOL. XXXVIII 
MARCH, 1931 
No. 1 
THE GENITALIA AND TERMINAL STRUCTURES OF 
THE MALE OF THE ARCHAIC MECOPTERON, 
NOTIOTHAUMA REEDI , COMPARED WITH 
RELATED HOLOMETABOLA FROM THE 
STANDPOINT OF PHYLOGENY 
By G. C. Crampton 
Massachusetts Agricultural College, Amherst, Mass. 
For the rare privilege of studying the hitherto unde- 
scribed male of the intensely interesting archaic Mecopteron 
Notiothauma reedi, McL. (which is in some respects a 
“living fossil” — the last remnant of a primitive relict fauna, 
representing as closely as any living form, the type an- 
cestral to the rest of the Mecoptera) I am deeply indebted 
to Professor Dr. C. E. Porter, who has done so much to 
advance the study of Entomology in Chile, and to Dr. E. P. 
Reed, the distinguished Chilean surgeon, who has been 
more than generous in supplying me with a wealth of ma- 
terial for morphological study. To both of these gentlemen 
I would express my profound gratitude for their aid in 
enabling me to carry out this investigation. 
The abdominal structures of the male of Notiothauma 
are the most remarkable and primitive to be found within 
the order Mecoptera, and could readily serve as the arche- 
types of these structures for the rest of the order. The 
primitive character of these structures is thus in harmony 
with the evidence of the archaic character of Notiothauma 
indicated by other features such as the wings, which have 
been discussed in Psyche, Vol. 87, p. 83, for 1930, where 
