Trelease.] 
434 
[March 15, 
the style comes with it. The stigma is confined to the lower sur- 
face of its tapering end, which in undisturbed flowers is curved 
upward so as to present the papillose face to the mouth of the 
corolla, fig. 46. 
Nectar is secreted by a large orange gland below the ovary, 
fig. 49. Fine hairs, glandular in part, are found on the calyx, 
the outside of the corolla, the ovary and the upper side of the 
style, figs. 49 and 52. Longer hairs are present on the filaments 
and the veins from which they spring, figs. 50-51, as well as on 
the ridges of the corolla which hold the style in place and across 
its floor for some distance near its mouth, figs. 46-48. These are 
evidently to be regarded as protections for the nectar against 
unbidden guests, and the rain and dew, though, as it must be 
said is the case in other plants, the presence of occasional ants 
in the flowers shows that the nectar guard is not entirely unsur- 
mountable. 
Not only are the flowers similar to those of G. isophylla, as 
described by Morren (2), in general form and size, but also in 
most details, the only differences that are noticeable being in the 
longer stamens and the style ; the former are unequal in G. iso- 
phylla, equal in G. anisopliylla ; while the latter does not show so 
marked an elbow at the base of the stigma in G. isophylla as in 
G. anisopliylla. Perhaps, too, the long hairs on the floor of the 
corolla do not extend so far forward in the latter as in the 
former species. 
The size and form of the flowers point to large bees like Bom- 
bus as the probable agents in fertilizing them. Here the area of 
pollination is removed from above to below the axis of the flow- 
ers, so that the pollen is carried on the under side of the insect, 
and not on its back as in Cystacanthus, the form of curvature of 
the corolla ensuring this contact. 
So far as could be seen the flowers are not dichogamous. It 
is evident, therefore, that their structure as thus far explained is 
conducive to close-fertilization, unless foreign pollen should prove 
prepotent ; for the pollen-covered part of an insect’s body must 
be brought in contact with the stigma, in its retreat from the 
the flower ; and although the concave face, which would be first 
met, is not stigmatic, the bend in the style is so great that at 
least the apical part of the stigma must be touched. 
