SIWALIK MAMMALIA. 
9 
figures, the fossil agrees with the kudu in all the above-mentioned respects, and 
also in the contour of the characteristic facial profile, and the position of the horn- 
core relatively to the orbit. The position of the large lachrymal fissure is still 
indicated in the fossil. The horn-cores are proportionately somewhat smaller than 
in the kudu, but their direction may have been approximately similar. This 
smaller size may be partly due to immaturity, but may also indicate a resemblance 
to & imberbis. The upper true molars of the existing species of Strepsiceros are 
relatively broad, the outer surface of each lobe being wide and without a distinct 
median costa ; the enamel is rugose, and there is no accessory internal column in 
the median valley. The teeth of the fossil agree very closely with those of the 
kudu, with the exception that they possess a distinct internal accessory column, 1 2 
but it is very doubtful if this difference necessarily implies generic distinction. An 
upper molar of a Siwalik ruminant has been described and figured in vol. III. p. 114. 
pi. XIII. figs. 9, 10, which it was suggested might possibly belong to the Pikermi 
genus Palceoryx ; since, however, this tooth appears to agree precisely with the 
molars of the present form it is probable that it should be referred to it : the tooth 
presents the characteristic antelopine (as opposed to cervine) feature of the accessory 
column being attached entirely to the posterior crescent. When adult it is probable 
that the fossil species was nearly as large as the kudu. 
Affinities. — The resemblance of the cranium under consideration to that of the 
existing Strepsiceros is so close that it may safely be regarded as indicating an allied, 
and probably ancestral form, which may be at least provisionally included in the 
same genus. This form is certainly specifically distinct from the two existing 
species, and is larger and otherwise different from Trayelaphus, and therefore 
probably also from Protragelaphus (of which the skull is not figured) ; it is moreover 
widely different from Palaeoreas? 
Distribution. — If the molar figured in vol. III. pi. XIII. figs. 9, 10, belong to 
the present form, its distribution will have extended from Perim Island to the 
Punjab. 
Genus. BOSELAPHUS, Blainville. 3 
Syn. P or tax, Hamilton Smith. 
The genus is confined to India, where it has existed since the Siwalik epoch. 
Boselaphus, sp. nobis.* 
Summary. — This form is known by teeth from the Siwaliks of the Punjab 
described and figured in vol. III. pp. 114-116. pi. XIII. which with the exception of 
their superior size are indistinguishable from those of B. trayocamelus. The upper 
1 The first true molar of one side has been cleaned in order to exhibit this structure ; the accessory column was 
unfortunately cut through, hut its section is visible. 
2 Compare Gaudry, “Les Enchainements, &c. — Mamra. Tert.” p. 82. fig. 91. 
3 ‘ Bull. Soc. Philom. Paris.’ 1816. p. 75. 
4 Supra vol. III. p. 114 (1884). 
C 
