426 
Fishery Bulletin 115(3) 
Table 2 Continued 
Scientific name 
2012 
2013 
2014 
2015 
Overall mean 
Holacanthus ciliaris 
0.08 
0.05 
0.11 
0.12 
0.09 
Cephalopholis cruentata 
0.14 
0.00 
0.09 
0.08 
0.08 
Holocentrus adscensionis 
0.11 
0.11 
0.00 
0.00 
0.06 
Seriola rivoliana 
0.03 
0.08 
0.09 
0.00 
0.05 
Halichoeres bathyphilus 
0.00 
0.00 
0.14 
0.04 
0.05 
Muraenidae 
0.01 
0.05 
0.03 
0.08 
0.04 
Pomacanthus paru 
0.09 
0.00 
0.03 
0.04 
0.04 
Rypticus saponaceus 
0.10 
0.03 
0.00 
0.02 
0.04 
Mycteroperca interstitialis 
0.11 
0.03 
0.00 
0.00 
0.03 
Mycteroperca bonaci 
0.06 
0.00 
0.00 
0.08 
0.03 
Pseudupeneus maculatus 
0.03 
0.00 
0.11 
0.00 
0.03 
Acanthurus spp. 
0.00 
0.03 
0.09 
0.02 
0.03 
Pomacanthus arcuatus 
0.00 
0.05 
0.03 
0.04 
0.03 
Haemulon album 
0.11 
0.00 
0.00 
0.00 
0.03 
Gymnothorax moringa 
0.03 
0.00 
0.09 
0.00 
0.03 
Myripristis jacobus 
0.10 
0.00 
0.00 
0.00 
0.03 
Aulostomus maculatus 
0.01 
0.00 
0.03 
0.04 
0.02 
Epinephelus adscensionis 
0.01 
0.00 
0.00 
0.06 
0.02 
Balistes vetula 
0.01 
0.03 
0.03 
0.00 
0.02 
Epinephelus guttatus 
0.03 
0.00 
0.00 
0.04 
0.02 
Haemulon melanurum 
0.00 
0.05 
0.00 
0.00 
0.01 
Lutjanus analis 
0.05 
0.00 
0.00 
0.00 
0.01 
Priacanthus arenatus 
0.00 
0.03 
0.00 
0.02 
0.01 
Seriola dumerili 
0.01 
0.00 
0.03 
0.00 
0.01 
Lachnolaimus maximus 
0.00 
0.00 
0.00 
0.04 
0.01 
Seriola spp. 
0.00 
0.00 
0.00 
0.04 
0.01 
Acanthurus coeruleus 
0.00 
0.00 
0.03 
0.00 
0.01 
Lutjanus buccanella 
0.00 
0.00 
0.03 
0.00 
0.01 
Lutjanus campechanus 
0.00 
0.00 
0.03 
0.00 
0.01 
Lutjanus griseus 
0.00 
0.00 
0.03 
0.00 
0.01 
Monacanthus tuckeri 
0.00 
0.00 
0.03 
0.00 
0.01 
Opsanus spp. 
0.00 
0.00 
0.03 
0.00 
0.01 
Serranus phoebe 
0.00 
0.00 
0.03 
0.00 
0.01 
Epinephelus itajara 
0.00 
0.03 
0.00 
0.00 
0.01 
Haemulon spp. 
0.00 
0.03 
0.00 
0.00 
0.01 
Balistes spp. 
0.03 
0.00 
0.00 
0.00 
0.01 
Neoniphon marianus 
0.03 
0.00 
0.00 
0.00 
0.01 
Balistes capriscus 
0.00 
0.00 
0.00 
0.02 
0.00 
Paranthias furcifer 
0.00 
0.00 
0.00 
0.02 
0.00 
Serranus annularis 
0.00 
0.00 
0.00 
0.02 
0.00 
Mycteroperca venenosa 
0.01 
0.00 
0.00 
0.00 
0.00 
Scarus coelestinus 
0.01 
0.00 
0.00 
0.00 
0.00 
used the grouper pits, cardinalfish, damselfish, and 
anthiids were the most abundant taxa (mean: 54.70 
individuals/pit [SE 6.70]). The red grouper and scamp 
(. Mycteroperca phenax ) were the most abundant eco- 
nomically important species (mean: 0.40 individuals/pit 
[SE 0.04] for red grouper and 0.35 individuals/pit [SE 
0.07] for scamp). Average abundance of unidentifiable 
fish was 13.91 individuals/pit (SE 3.60). 
The percentage of grouper pits with red grouper in 
them was 37.5% in 2012, 46.3% in 2013, 45.7% in 2014, 
and 34.6% in 2015 — proportions that were not signifi- 
cantly different (one-way ANOVA: P=0.61). There were 
never multiple red groupers in any one pit, and they 
were distributed evenly inside and outside the HAPC, 
as well as across the various regions. Frequency of oc- 
currence for red grouper was 40.6% inside and 35.1% 
outside the HAPC, proportions that were not signifi- 
cantly different (one-way ANOVA: P=0.54). Their fre- 
quency of occurrence was 40.4% on the main ridge, 
37.5% at off main ridge, 37.5% in the central basin, 
and 44% on the west ridge. These values were also not 
significantly different (one-way ANOVA: P=0.97). Of 
red grouper that could be measured from the lasers 
mounted on the ROV, total length ranged from 50 to 80 
cm (average: 60 cm). 
In contrast, lionfish were observed in 72.5% of grou- 
