MacDoitgal. — Symbiotic Saprophytism . 1 7 
internodes, bearing longer and subdivided branches. The 
third offshoot branched near the point of origin, and the 
branches were subdivided in such manner as to form an almost 
solid mass of coralloid formations. Two plants arose from 
different branches of this offshoot. One of the plants from 
this offshoot was detached with a mass of the coralloid stem 
for anatomical examination, and the remainder were photo- 
graphed and placed in a cold house for three months, where 
they perished by freezing. It was most noticeable that the 
formation of young plants had not greatly decreased the size 
of the parent corm. In this instance the growth of the 
offshoot had consumed the reserve material in the lower 
part of the corm ; but if the upper part had undergone 
the usual shrinkage, it had subsequently regained its original 
plumpness. The formation of the great mass of coralloid 
offshoots not only did not involve the expenditure of the 
amount of food usually consumed by one plant, but was 
accompanied by an increase of the material in the parent 
corm, indicating a distinct change in the nutritive conditions. 
The coralloid shoot differs chiefly from the type by its 
profuse branching and the enormous multiplication of the 
internodes, with a corresponding increase of surface. The 
extension of surface is carried still further by the formation 
of great numbers of trichomes, which replace the scale-leaves. 
The latter appear to be almost wholly lacking. The clumps 
of trichomes are so closely crowded as to give it the appear- 
ance of being covered with root-hairs to which the particles 
of humus cling, and are most numerous near the apices of 
the branches. The epidermis is composed of irregular 
flattened cells, with walls strongly thickened. The stomata 
are more numerous and larger than in normal offshoots. 
An ectotropic Fungus covers the surface in patches in much 
greater profusion than in the normal offshoot, and penetrates 
the epidermal cells in the same manner as in roots. The 
hypoderm consists of ovate cylindrical elements with thickened 
walls. The two layers are rich in protoplasm and are 
separated by intercellular spaces. The trichomes are large 
C 
