536 WorsdelL — Observations on the Vascular 
Abietineae. 
Pinus sylvestris , L. 
In the female cone of this plant, four bundles leave the 
central cylinder of the axis of the cone : a lower one with 
normal orientation, an upper one with inverted orientation 
(which arises by fusion of two bundles springing right and 
left from distinct strands of the cylinder), and two lateral 
ones orientated sideways (Fig. 9). These all pass through 
the cortex and enter the base of the appendage, without 
undergoing further division or alteration in position. The 
lower bundle enters the small bract which is adnate for 
a short distance to the much larger seminiferous scale. The 
three other bundles enter the latter, the two lateral moving 
up so as to form a single row with the upper bundle (Fig. 10); 
by subsequent division a considerable number of bundles is 
formed. The plant differs from Araucaria in the fact that 
four bundles, instead of two, leave the cylinder of the axis, 
the bundles which supply the seminiferous scale and bract 
respectively thus arising independently from the cylinder of the 
axis , and no connexions occurring between the two systems 
during their course through the cortex. But in Araucaria 
Bidwillii , Hook., as we have seen, the two systems also arise 
independently from the central cylinder of the axis, and they 
appear to preserve their independence during their subsequent 
course far more than is the case in A. Cookii , R. Br. ; A. Bid- 
willii , Hook., however, would appear to be an exception to 
the general rule prevailing in that genus. The direct 
explanation of the structure in Pinus sylvestris , L., lies in the 
fact of the more complete separation of the bract and semini- 
ferous scale than is the case in Araucaria , the almost complete 
fusion in the latter of the two foliar organs having occasioned 
a similar fusion between the two otherwise distinct vascular 
systems, and the latter fact not necessarily implying that the 
appendage in Araucaria is really a single organ. 
A single bundle with normal orientation, leaves the cylinder 
