584 Wager. — The Sexuality of the Fungi . 
evidence strongly in favour of regarding these organs as sexual, 
but that it supports Harper’s conclusions as to fertilization. 
Dangeard \ in fact, himself, in speaking of his own view of the 
sexuality of Sphaerotheca , says : — - Cette fecondation (nuclear 
fusion in the ascus) laisse meme toute liberte d’interpretation 
au sujet des archicarpes et des branches antheridiennes ; elle 
exige simplement que ces organes soient devenus inutiles ; 
s’ils remplissent encore leur fonction, s’il y a fertilisation, 
basque doit en provenir directement ; et c’est bien le cas, ainsi 
qu’en temoignent les Eremascus Eidam et les Dipodascusl 
It is of course possible that fertilization may not always 
take place in individuals of this species, which would account 
to some extent for the discrepancy between the observations 
of these two authors ; and in this connexion the observations 
made by Mary M. Nichols 2 on certain pyrenomycetous Fungi, 
which distinctly support the views maintained by Harper, are 
of considerable interest. The author states that in Ceratostoma 
brevirostre and Hypocopra the antheridia fuse with the archi- 
carps in some cases, whilst in others the archicarps appear to 
develop without fertilization. In Teichosporella there remains 
only a possible rudiment of an antheridium, while in Teicho- 
spora this has often entirely disappeared. 
In the Ustilagineae Dangeard 3 states that a fusion of two 
nuclei takes place in the young resting spores (brand spores) 
of Doassansia Alismatis and Entyloma Glaucii , Dang. Doas- 
sansia Alismatis forms brown pustules on the leaves of Alisma 
Plantago , and its slender mycelium is found in abundance in 
the intercellular spaces of the mesophyll. Each cell contains 
several nuclei. In Entyloma Glaucii 4 also the mycelium 
is found abundantly in the intercellular spaces of the host- 
plant, and the cells are multinucleate. In both species the 
1 Dangeard, loc. cit., p. 252. 
2 Mary M. Nichols, The Morphology and Development of certain pyreno- 
mycetous Fungi. Bot. Gazette, vol. xxii, 1896, p. 301. 
3 Dangeard, Recherches sur la Reproduction sexuelle des Champignons. Le 
Botaniste, iii, 1893-4, p. 221. 
4 Dangeard, La Reproduction sexuelle de /’ Entyloma Glaucii (Dang.). Le 
Botaniste, iv, 1894-5, p. 12. 
