92 Campbell ’ — Sttidies on some Javanese Anthocerotaccae. II. 
three trees near the edge of the forest, there was found growing a small 
species (PL IX, Fig. 4), which was assumed to be D. javanicus. Although 
careful search was made in various localities, no other specimens were 
found. A second, much larger species (Fig. 1) was subsequently met with 
nearer the summit of Mount Gedeh. This was only found once, but was 
growing abundantly on the twigs of an undetermined shrub, and was quite 
conspicuous, while the other species growing among the mosses and liver- 
worts on the bark of trees was very hard to find. The large species was 
growing at an elevation of 2,200 metres, which was rather unexpected, as 
Dendroceros usually prefers the warmer regions nearer sea-level. 
The determination of the two species must be left until a comparison 
can be made with authentic specimens of D. javanicus. We shall simply 
refer to them here as species A, the large form from the higher station, and 
species B, the smaller form from Tjibodas. The former species is very 
much larger than the other (see PI. IX, Figs. 1 and 4), and in many respects 
seems to agree with the plant which Leitgeb studied under the name 
D. javanicus} The magnification of Leitgeb’s figures is not indicated, and 
it is impossible therefore to judge of the size of the plant studied by him. 
The thallus of form A (Fig. 1) is about 3 cm. in length, and more or less 
regularly pinnately branched. The midrib, as in all species of Dendroceros , 
is sharply defined, and the wings of the thallus consist of a thin lamina 
composed of but a single layer of cells. This lamina in the species in 
question is very much lobed and folded, so that it closely resembles such a 
liverwort as Fossoinbronia , the lobes of the lamina simulating leaves. In 
this respect it resembles Leitgeb’s figure of D. javanicus , but the margin is 
even more irregular than in his figure. It also resembles the West Indian 
D. crispus ) but is very much larger than any specimens of that species 
collected by the writer in Jamaica, and the margin of the thallus is very 
much more lobed and folded. The cells of the lamina are destitute of the 
large lacunae which are very conspicuous in the Tjibodas plant (Fig. 6), and 
in this respect it differs from Leitgeb’s plant, which is described as having 
large lacunae in the lamina. The cells of the lamina show the collenchyma- 
like thickenings at the angles that Leitgeb describes in Dendroceros 
Breutelii. The midrib is very massive, and contains large intercellular 
spaces, but these are not air-chambers, as Leitgeb states. He compares 
them to the air-chamber of the Marchantiaceae, but they are filled with 
mucilage, as they are in other Anthocerotaceae where such spaces are 
present. Leitgeb was probably misled by having to deal with dried 
material, where the drying up of the mucilage or its colourless character 
may have made him overlook the contents of these spaces. In stained 
microtome sections it is very evident that these intercellular spaces are not 
air-chambers. 
1 Leitgeb, loc. cit, Plate II, Fig. 20. 
