94 Campbell. — Studies on some Javanese Anthocerotaceae . II. 
Anthoceros , but this is not very marked. As in other Anthocerotaceae, 
the archegonium projects but slightly above the level of the thallus. There 
are from two to four cover-cells in the ripe archegonium (Fig. 13), and these 
are thrown off when the archegonium opens. The neck-canal-cells usually 
number five. 
The antheridia (Figs. 14 to 22), as in other species of Dendroceros , are 
borne singly in the antheridial cavity. No attempt was made to follow out 
the early development, as there was no indication that this differed in any 
marked degree from that of the other species that had been studied. There 
is no doubt of the endogenous origin of the antheridium, and, as in all 
other described species of Dendroceros , the archegonium lies nearly 
horizontal in the antheridial chamber and develops an extremely long 
pedicel, which is coiled up in the cavity. Leitgeb supposes that at 
maturity the stalk straightens out and projects the archegonium above the 
surface of the thallus. This point, however, was not investigated. The 
body of the antheridium is oval, and very early in its history the single 
layer of peripheral cells is separated from the inner spermatogenic tissue. 
Cross-sections of the pedicel (Fig. 19,/) show it to be composed of two rows 
of cells in all the specimens examined in both species. This agrees with 
Leitgeb’s account of the species studied by him, but in D. Breutelii there 
may be four rows in some cases. 1 
The spermatozoids are too small to make a study of the spermato- 
genesis satisfactory, and no attempt was made to trace the development 
of the spermatozoids. After the last mitosis the spermatocytes are in 
pairs, apparently not separated by a cellulose membrane (Fig. 18). This 
is much like what occurs in many Hepaticae. Presumably the cilia arise 
from a blepharoplast, but this point was not demonstrated. The mature 
antheridium is very large in the species A (Fig. 19), being about 350 /x in 
diameter. This is more than twice the size Leitgeb gives for D. javanicus 
(loc. cit. , p. 32). The antheridium in the second species is much smaller 
(Fig. 22). 
The Embryo. 
Leitgeb describes the older embryos of Dendroceros, but did not 
obtain the earlier stages. The writer succeeded in obtaining younger 
stages of D. Breutelii (loc. cit.), but the first divisions were not seen, 
although it was conjectured that the first division was a longitudinal 
one, as in Anthoceros. This has proved to be also the case in the two 
species under consideration (Fig. 23), and it will be safe to assume that this 
is the rule in Dendroceros. The second walls are transverse and nearly 
median. This first transverse division probably determines the boundary 
between the foot and the capsule. The next divisions are vertical, but 
Campbell, loc. cit.. p. 472. 
