g6 Campbell. — Studies on some Javanese Anthocerotaceae . II. 
recognized by their rounded form and greater diameter. The sterile cells 
remain narrow and are soon easily distinguishable from the rounded 
spore-mother-cells. The division of the latter was not followed, as there 
was no evidence that it was different from the other forms that have 
been studied. The two Javanese species do not differ much in the size 
of the ripe spores (Fig. 30), but the species A has both the columella 
and the outer tissue much more developed than the smaller one. In the 
latter there are seldom more than four layers of cells in the sporogonium- 
wall, while in the larger species there may be as many as eight. The 
superficial cells of the smaller species are longer and narrower than in 
the larger one, and in the latter they often project slightly. 
After the division of the spore-mother-cell is completed, the spores 
remain together in tetrads until they are almost ready to be discharged 
from the sporogonium. While still together, a thin, finely papillate 
perinium is developed and the spores undergo repeated division, so that 
each one is a multicellular body of considerable size (Fig. 30). Both 
of the Javanese species agree in this respect, and this is known to be 
the case also in a number of others, e. g. D. crispus , D. cichoraceus. 
The cell-mass derived from the spores has a large amount of chlorophyll, 
and really germination begins inside the sporogonium, as it does in 
Pellia and Fegatella. 
The elaters, as in the other species of Dendroceros , are multicellular, 
and are to be considered as fragments of the net of sterile cells which 
incloses the spores. There is a broad spiral band of a yellowish brown 
colour, which is scarcely interrupted by the division walls between the 
cells of the elater (Fig. 31). 
NOTOTHYLAS JAVAN ICUS. 
In the garden at Buitenzorg,and at other places in the vicinity, a species 
of Notothylas , presumably N. javanicus , was not at all uncommon. 
Material of this species was collected in order to compare it with the 
common American species, N. orbicidaris , which is the only species 
that has been completely investigated. 1 The statement of Leitgeb, that 
the columella in Notothylas is not infrequently quite absent or may be 
of secondary origin, was not confirmed by Mottier in the case of 
N. orbicidaris. N. javanicus was examined carefully with reference to 
the same point, and the conclusions reached agree essentially with those 
of Mottier with reference to N. orbicidaris. N. javanicus closely resembles 
in general appearance N. orbicidaris. The thallus is usually nearly orbicular, 
due to a repeated and rapid dichotomy. It is somewhat yellowish-green 
in colour, and the sporogonia project but little from the involucre. Where 
1 Mottier, Contributions to the Life History of Notothylas , Ann. of Bot., viii, 1894 ; Campbell, 
Development of the Mosses and Ferns, First Ed., 1895. 
