Campbell. — Studies on some Javanese A nthocerotaceae. II. 97 
the sporogonia are abundant the short, nearly horizontal capsules occur 
quite thickly set round the margin of the thallus. Quite often dwarf 
sporogonia are met with, resembling those described in other species, 
and like these are probably due to imperfect nutrition. 
Sections of the thallus of N. javanicus show it to be quite solid, 
and no trace of the mucilage-cavities which are conspicuous in N. 
orbicularis can be found. The apical growth (PI. X, Pigs. 32, 33) is 
exactly like that of Anthoceros and Megaceros , there being two sets of 
segments, dorsal and ventral, cut off from the large initial cells. 
The Reproductive Organs. 
The archegonia are similar in origin and development to those of 
Anthcceros , but they are somewhat broader in the early stages, and the 
number of neck-canal-cells is usually less. In all the specimens of 
N. javanicus that were examined there were but three neck-canal-cells 
(Figs. 32, 34), and Mottier found this to be true in N. orbicularis , although 
the writer found as many as five in the latter species. The number of 
ripe archegonia that were examined in N. javanicus was not very large, 
and it is not impossible that in this species also there may exceptionally 
be more than three neck-canal-cells. 
In some cases the neck-cells are cut off by a longitudinal wall after 
the first transverse cell of the young archegonium has been formed (PI. X, 
Fig- 33)- In such cases the neck-canal-cells are narrower than when 
this longitudinal wall is suppressed. An abnormal case was seen in 
which the egg-cell had divided into two by a longitudinal division 
wall after the cutting off of the ventral canal-cell (P'ig. 34). There are 
usually two cap-cells present. Whether more than this may be formed 
in N. javanicus was not determined. The antheridia (Fig. 35) are 
formed usually four together in the antheridial cavity, and apparently 
do not differ in any important particular from those of N. orbicularis . 1 
The Embryo. 
Neither Leitgeb nor Mottier described the earliest stages of the 
embryo. The latter thinks that in N. orbicularis the first division-wall 
is transverse. This does not seem to be the case in N. javanicus to 
judge from the few young embryos that were examined. In one case a 
two-celled embryo was sectioned (Fig. 36). This was divided longi- 
tudinally exactly as in Anthoceros. The next youngest embryo seen 
(Fig. 37) was divided into three tiers by two transverse sets of walls, 
and in this case at least it was clear that the development of the columella 
and archesporium was confined to the uppermost segment of the embryo 
1 See Campbell, Mosses and Ferns, Second Ed., Fig. 80. 
